272 



UNITED STATES NATIONAL MUSEUM BULLETIN 207 



Euzophera, easily distinguished from that genus by the 

 close association of veins 3, 4 and 5 of forewing, and the 

 short ceU of hind wing. Contains one tropical Ameri- 

 can species. Hampson includes a species from Mada- 

 gascar but this is probably improperly placed. 



562. Edulica compedella (Zcller) 

 Figures 58, 585, 1071 



Euzophera compedella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 



p. 224, 1881. 

 Edulica compedella (Zeller) Ragonot, Monograph, pt. 2, p. ix, 



1901. — Hampson, in Ragonot, Monograph, pt. 2, p. 122, 



1901. 



Ground color of forewing varying from somber brown 

 to gray-brown; costal half of wing dusted with ashy 

 grayish white; the discal spots blackish, confluent; some 

 scattered blackish dots on several of the veins; ante- 

 medial line not defined; the subterminal one weakly so 

 by its dark inner and outer bordering lines, irregularly 

 dentate; terminal dots distinct, blackish. Hind wing 

 of male whitish with dark brown shading on veins and 

 along termen; of female smoky fuscous, darkening 

 towards apex and termen. Alar expanse, 20-25 mm. 



GenitaHa as given for the genus. 



Type localitt : Honda, Colombia (type in BM) . 



Food plant: Unknown. 



Distribution: Panama: Porto Bello (Apr., May). 

 Colombia: Honda. French Guiana: St. Jean 

 Maroni. Brazil: Amazonas, Ponte Nova (Rio 

 Xingu). 



Genera 163 and 164: Euzophera and Exuperius 



[Venational division A. Forewing with 11 veins; 4 and 5 stalked. 

 Hind wing with vein 2 from well before lower outer angle of cell; 

 3 and 5 connate; cell long; discocellular vein curved. Male geni- 

 talia with apical process of gnathos a stout, elongate hook with 

 pointed apex; transtiUa complete. Labial palpi upturned.] 



163, Genus Euzophera Zeller 



Stenoptycha Heinemann (not Zeller), Schmetterlinge Deutsch- 

 lands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 190, 1866. 



Melia Heinemann, Schmetterlinge Deutschlands und der 

 Schweiz, Abt. 2, vol. 1, pt. 2, p. 209, 1866 (new name, pre- 

 occupied, proposed for Stenoptycha Heinemann). 



Euzophera Zeller, Trans. Ent. Soc. London, ser. 3, vol. 5, p. 456, 

 1867 (new name for Stenoptycha Heinemann) ; Stettiner 

 Ent. Zeit., vol. 28, p. 377, 1867.— Hulst, Phycitidae of 

 N. Amer., p. 174, 1890 (cites type as Myelois cinerosella 

 Zeller). — Hampson, Moths, vol. 4, p. 72, 1896, in Blanford, 

 Fauna of British India (cites type as Ephestia biviella 

 Zeller); in Ragonot, Monograph, pt. 2, p. 36, 1901. — 

 Ragonot, Monograph, pt. 2, p. vii, 1901 (cites type as 

 Phycis pinguis Haworth). — Forbes, Cornell Mem. 68, p. 

 630, 1923. — Bisset, in Pierce and Metcalf, Genitalia of the 

 British Pyrales, p. 59, 1938 (notes fixation of type by 

 Hulst). — Janse, Journ. Ent. Soc. South Africa, vol. 8, p. 31, 

 1945. (Type of genus: Myelois cinerosella Zeller; figs. 77, 

 578, 1063.) 



Tongue well developed. Antenna simple, shaft 

 pubescent (ciliations very short). Labial palpus up- 

 turned, reaching to or almost to vertex; second segment 

 rough scaled; third segment about half as long as 



second, acuminate. Maxillary palpus subsquamous 

 (somewhat rough scaled), appressed to face. Forewing 

 smooth; 11 veins; vein 2 from well before lower, outer 

 angle of cell; 3 from the angle; 4 and 5 stalked (for less 

 than half the lengths) ; 6 from below upper angle of cell, 

 straight; 8 and 9 long stalked; 10 normally from the 

 cell, separated from or approximate to (rarely connate 

 with or from the stalk of) 8-9; male without costal fold. 

 Hind wing with vein 2 from well before lower outer 

 angle of cell; 3 and 5 from the angle, connate, rarely (in 

 individual specimens) very shortly stalked; 7 and 8 

 strongly anastomosed beyond cell (for more than haK 

 their lengths); ceU long, slightly more than half the 

 length of wing; discocellular vein curved. Eighth 

 abdominal segment of male simple except in cinerosella 

 which has a pair of short ventrolateral hair tufts. 



Male genitalia with apical process of gnathos devel- 

 oped as an elongate, stout, simple hook with pointed 

 apex. Uncus stout, broadly triangulate. Transtilla 

 complete, developed as a strongly sclerotized arch with 

 prominent, caudally projecting, lateral lobes. Harpe of 

 nearly equal width throughout; terminal margin 

 rounded and with a short rounded projection from apex 

 of costa; costa otherwise, for most of its length, strongly 

 sclerotized; clasper not developed. Anellus a slightly 

 curved U-shaped shield, the lateral lobes well developed 

 and strongly sclerotized in the American species. 

 Aedeagus moderately long, stout; penis armed with 

 numerous, short, sharp spines. Vinculum stout, as long 

 as (or very little longer than) broad; terminal margin 

 rounded. 



Female genitalia with numerous sclerotized or scob- 

 inate folds in bursa copulatrix adjacent to or at junction 

 with ductus bursae (except in the European type, 

 cinerosella), finely scobinate over most of inner surface; 

 signum present as a small, cupped, scobinate plate; 

 ductus bursae with genital opening simple; ductus 

 seminalis from bursa near signum (in American species) 

 or from terminal end of bursa (in cinerosella). Dorsal 

 sclerotized area of eighth segment coUar a narrow band 

 with central tonguelike projection. 



Euzophera, like Zophodia, has served as the receptacle 

 for a heterogeneous lot of species, most of which must 

 be referred to various other genera. As here defined 

 the genus contains only three described American 

 species that can be referred to it with any certainty. 

 These form a compact group with constant genitalic 

 characters and similar forewing pattern: antemedian 

 line far out on wing, subterminal line well back from the 

 terminal margin, the space between the lines corre- 

 spondingly reduced, and a white spot on discocellular 

 vein. On venation, palpi, and male genitalia they 

 agree with the European type of genus, cinerosella, but 

 differ from it in wing pattern and two details of struc- 

 ture, cinerosella having hair tufts on the eighth abdom- 

 inal segment of the male, and the ductus seminalis from 

 the anterior end of the female bursa (figs. 578b, 1063). 

 Eventually it may be possible to give our American 

 species a separate generic designation; but upon present 

 knowledge they must be retained in Euzophera. 



