366 U.S. NATIONAL MUSEUM BULLETIN 271 



Type-species: Tryphosella sarsi Bonnier, 1893 (present selection), 

 {—Tryphosa nana of Sars, 1895, not of Boeck, 1861). Bonnier in- 

 tended the name Tryphosella to replace Sars' usage of the name 

 Tryphosa for a group of species distinct from Orchomenella Sars 

 (=Orchomene Boeck). Sars did not realize that those species he called 

 Orchomenella were congeneric with Tryphosa. Stebbing (1906) also 

 failed to recognize Bonnier's perception and restricted Tryphosella to 

 a single species, Tryphosa barbatipes Stebbing, 1888, which later has 

 been considered to be congeneric with Uristes Dana. Thus J. L. 

 Barnard's (1962d) move of Tryphosella to C/m^es was illegal because 

 the type-species of Tryphosella had never been selected. Legaliza- 

 tion of Bonnier's intent by selection of the type-species that he 

 obviously w^ould choose through his comments and his listing of it 

 in primary position now permits reinstatement of a "tryphosa" 

 group to be called a "tryphosella" group of species. They are close 

 to Tmetonyx Stebbing, 1906, but have coxa 1 distinctly tapering 

 dis tally. Lepidepecreopsis Stephensen may be a synonym of Tryphosella 

 as Gurjanova (1951) removed the former to Tryphosa. See 

 discussion with Tmetonyx. 



Epistome broadly rounded and dominating upper lip, both sepa- 

 rated by a notch; mandibular molar subcolumnar or cuboid, weakly 

 setulose and poorly ridged if at all (Lepidepecreopsis), palp attached 

 level with molar; coxa 1 slightly shortened, distinctly tapering 

 (Anonyx, Tmetonyx) and partially hidden by coxa 2; gnathopod 1 

 subchelate, articles 5 and 6 subequal in length; telson of medium 

 length, deeply cleft; head oj normal size (Uristes); (dactyl of gnathopod 

 1 with or without inner tooth). Species: ca. 63, cosmopolitan cold- 

 water, littoral to hadal, in low latitudes primarily bathyal-abyssal. 

 Composition: all species formerly known under "Tryphosa" and 

 "Tmetonyx" since 1895 except their type-species. 



Tryphosites Sars 



Tryphosites Sars, 1895. — Stebbing, 1906. 



Type-species: Anonyx longipes Bate and Westwood, 1863 (orig- 

 inal designation). 



Epistome acutely produced in front of upper lip, upper lip also 

 slightly lobate; mandibular molar ridged, palp attached level with 

 molar; gnathopod 1 subchelate, article 5 longer than 6; inner ramus 

 of uropod 2 constricted; telson of medium length, deeply cleft. Species: 

 5, bipolar, coldwater submergent, littoral to abyssal. 



