MARESPEi GAMMJARIDEAN AMPHIPODA 467 



There are exceptions to the neat morphological partitioning of 

 talitrid-like animals but there are definitely three or more facies and 

 perhaps three lines of evolution as Bulycheva's arrangement points 

 out. She did not assign all existing genera to families, which I do in 

 the following scheme, considering for the uses of this handbook that 

 they are subfamilies. 



Subfamily Talitrinae: Talitrus, Talorchestia, Orchestia, Orchestoidea. 

 Subfamily Hyalellinae: Hyalella, Parhyalella, Insula, Chiltonia, 



Afrochiltonia, Neohule, Austrochiltonia, [Najnaf] but see below. 

 Subfamily Hyalinae: Hyale, Parhyale, Allorchestes , Parallorchestes. 

 ? (Subfamily Najninae) : Najna. 

 ?(Subfamily Dogielienotinae) : See as a family, p. 207. 



Probably Neohule Haswell was improperly described and classified. 

 It has not been recovered since its original description in 1880. It 

 has the aspect of a Parhyalella but may belong to another family, 

 such as the Kuriidae, Prophliantidae, or Lysianassidae. Its "biramous 

 posterior pleopoda" (presumably uropoda) may be an erroneous 

 observation and the condition of the mandibular molar was not 

 stated. It is the only talitrid presumed to have clearly biramous third 

 uropods and so confuses what would otherwise be a clear diagnosis 

 of the family. 



Key to the Genera of Talitroidea 



Neohule Haswell, a dubious genus, is not included. 



1. Terminal maxillipedal palp article unguiform (claw-shaped) (figs. 169j,fc) . 2 

 Terminal maxillipedal palp article not unguiform (figs. lQ^l,m,n) .... 8 



2. Uropod 3 with minute inner ramus (fig. 169o) 3 



Uropod 3 uniramous (fig. 169g) 4 



3. Palp of maxilla 1 uniarticulate (fig. 169/i) Parhyale 



Palp of maxilla 1 biarticulate (often faintly so) (fig. 169g) . . Parallorchestes 



4. Telson entire (fig. 169r) 5 



Telson cleft (fig. 169s) 7 



5. Maxilla 1 lacking palp (fig. 169/) Parhyalella 



Maxilla 1 bearing palp (figs. \Q^g,h) 6 



6. Male gnathopods 1 and 2 similar to each other in size and morphology 



(fig. 169a) Najna and Insula 



Male gnathopod 2 much larger and of different morphology than gnathopod 

 1 (fig. 1686) Hyalella (freshwater) 



7. Article 5 of male gnathopod 2 masked by articles 4 and 6 (fig. 168e) . Hyale 

 Article 5 of male gnathopod 2 produced between articles 4 and 6 (fig. 



168gr) Allorchestes 



8. Male gnathopod 1 simple (fig. 168/i) 9 



Male gnathopod 1 subchelate (fig. 1680 10 



9. Male gnathopod 2 feebly chelate (fig. 168/) Talitrus (terrestrial) 



Male gnathopod 2 large and strongly subchelate (fig. 168e). 



Orchestoidea (terrestrial) 



