ART. 16 



FOSSIL REPTILES FROM MONTANA GILMORE 



31 



In the light of our present knowledge of the genus Nodosaurus, 

 I can see no justification for its inclusion in a separate subfamily, 

 the Nodosaurinae, as proposed by Nopsca. Certainly nothing is 

 found in its known anatomy that would prevent its assignment to 

 the Ankylosaurinae. 



The inclusion of the genus Trobdon in the Nodosauridae by Nopcsa 

 is so obviously in error that I take this opportunity to challenge the 

 propriety of the assignment. Troodon is a typically bipedal animal 

 with greatly reduced fore limbs, and a slender elongated scapula; 

 premaxillary teeth present ; ilium with postacetabular portion verti- 

 cal and subequal in length with preacetabular portion ; ischium slen- 

 der, ceratopsian like; ribs slender; tibia and femur subequal in 

 length; pes having phalanges with tongue and groove joints; a 

 segmented abdominal cuirass. 



The features briefly reviewed above are certainly an array of 

 characters that, taken together, do not permit of the inclusion of 

 the genus in the family Nodosauridae. 



At the time of describing the osteology of Troodon -^ attention 

 was directed to the curious mixture of characters found in the skull 

 and skeleton and it was my conclusion that the relationships of 

 Troodon would be best expressed by its reference to a distinct family 

 of the Ornithopoda known as the Troodontidae, a conclusion, after 

 again reviewing all of the evidence, that I see no reason to alter. 



Chronological list of North American Nodosauridae 



Name, authority, and date 



Palaeoscincus costatus Leidy, 1856 



Priconodon crassus Marsh, 1888 



Nodosaurus texiUus Marsh, 1889 



Palaeoscincus lutus Marsh, 1892 



HoplitosauTus marshi Lucas, 1901 



Palaeoscincus asper Lambe, 1902 



Sterocephalus tutus Lambe, 1902 



Stegopelta landerensis Willi^ton, 1905 



AnkylosauTUs magnivertus urovvn, 1SG8 



Hierosaurus sternbergi Wieland, 1909 



Panoplosaurus mirus Lambe, 1919 



Dyoplosaurus acutosquameus Parks, 1924. 



ScolosauTus cutleri Nopcsa, 1928 



Edmontonia longiceps Sternberg, 1928__. 

 Anodontosaurus lavibei Sternberg, 1929. 

 Palaeoscincus rugosidens Gilmore, 1830.. 



Formation and locality 



Judith River, Mont.. 



Arundel, Md 



Benton, Wyo 



Lance, Wyo 



Lakota, S. Dak 



Belly River, Alberta. 

 do 



Benton, Wyo 



Hell Creek, Mont. 



Niobrara, Kans 



Belly River, Alberta. 

 do 



Edmonton, Alberta, 

 .do. 



Nature of type 



Tvro Medicine, Mont... 



Tooth. 

 Do. 

 Parts of skeleton and armor. 

 Tooth. 



Parts of skeleton, dermal armor. 

 Tooth. 



Fragment of skull, dermal armor. 

 Parts of skeleton, dermal armor. 

 Skull; much of skeleton, andder- 



mal armor. ,_ 



Dermal armor. ''"' '" y^ 



Skull; parts of skeleton and armor. 

 Fragment of skull; posterior part of 



skeleton. ^i 



Much of armor and skeleton. *;.j 

 Skull, armor, and parts of skeleton. 

 Skull and armor. k4 



Skull; much of skeleton and armor. 



2. ON A SKULL OF THE GENUS DYOPLOSAURUS 

 Plate 9, F gures 1 and 2 



In 1924 Prof. W. A. Parks ^^ established the genus Dyoplosaurus, 

 based on a partial but very interesting skeleton, consisting princi- 



=8 Gilmore, Charlps W., Bull. No. 1, University of Alberta, 1924, p. 39. 

 2> Univ. Toronto Studies, No. 18, 1924, pp. 5-24. 



