BALFOUR AND SMITH—MOULTONIA. 355 
the lamina of the cotyledon at its end and that the inflorescence 
meristem instead of being strictly limited to an area at the base 
of the cotyledon, is spread along the hypocotyl and also along 
the lamina. 
As an intermediate condition we may bring into the case 
the evidence offered by Monophyllaea. 
We have pointed out that Moultonia is allied to Monophyllaea. 
This genus we only know from the description and figure given 
by Clarke.* Clarke suggests, we think rightly, that the folium 
unicum of his plant is a cotyledon. There appears to be in 
Monophyllaea a single stalked leaf—to speak conventionally— 
as in Moultonia, but the stalk is shorter and at the junction of 
lamina and stalk scapose inflorescences arise. At the summit 
of the scapes the flowers are disposed in a unilateral spicate 
raceme which curls over circinately. We should interpret all 
this construction as that of a hypocotyl elongated in the form 
of a petiole and bearing at its extremity a cotyledonary lamina. 
We should expect in the young protocorm to find a second 
smaller arrested cotyledon. At the point where cotyledonary 
lamina and hypocotyl join the inflorescence arises from the 
hypocotyl. The meristem for the inflorescence is localised at 
Of the allied Epithema we have insufficient knowledge to 
allow of our making a suggestion of interpretation. 
The following morphological series within the plants named 
issues from what we have said :— 
Streptocarpus polyanthus, Hook.—hypocotyl not elongated, 
cotyledon sessile, scapose inflorescence with open biparous 
branching developing from hypocotyl base of cotyledonar 
lamina. 
Siveptocarpus Rexii, Lindl.—hypocotyl shortly elongated, 
giving apparent stalk to cotyledon, scapose inflorescence 
(x flower) developing from hypocotyl at base of cotyle- 
donar lamina. 
Monophyllaea—hypocotyl much elongated, giving apparent, 
long stalk to cotyledon, scaposé inflorescence with 
unilateral racemes developing from hypotyl at base of 
cotyledonar lamina. tc 
Moultonia—hypocotyl very much elongated, giving apparent 
very long stalk to cotyledon, ‘inflorescence disposed in 
umbels originating along middle line on upper surface 
of whole length of elongated hypocotyl and on midrib of 
lamina (but not showing special relation to the primary 
veins). : 
* C, B. Clarke, Cyrtandreae in De Candolle Monogr. Phanerog. v (1883-87), 
181, t. xx. 
