440 E. Ray Lankester — Discovery of Sabre-toothed Tiger 



I believe that it is only to the part noticed in the last paragraph 

 that Mr. Weston's remarks refer. 



Postcript. — In the discussion of this paper, Sir C. Lyell said 

 that he had found in this raised beach more species of shells than 

 those above mentioned ; and Mr. Prestwich said that at one spot the 

 number of young shells was remarkably great as compared with that 

 of older shells. 



III. — On the Oocukrenoe of Machaikodus in the Forest-bed 



OF Norfolk. 



By E. Ray Lankester, B.A. Oxon. 



[PLATE XVI.] 



A SPECIAL interest has always attached itself in this country 

 to the remains of that rare and bizarre carnivor, the Sabre- 

 toothed tiger. For as yet it has only been indicated to us on this 

 side of the Channel by the most fragmentary and unique specimens 

 — the two teeth, incisor and canine, obtained by the Eev. Mr. 

 MacEnery, from Kent's Hole, Torquay, the authenticity of which 

 have even been doubted. Whilst, however, palaeontologists have 

 pretty generally come to the conclusion that these two isolated 

 teeth are really the remains of a Devon Sabre-tooth, no further 

 specimens have, I believe, been recorded from other caverns in this 

 country. 



The forest-bed of Norfolk, which furnishes remains of BMnoceros 

 Etruscus in some abundance, and of Ursus Arvernensis — and other 

 forms associated in central France with species of Machairodus — has 

 hitherto not given up any remains of that carnivor to the energetic 

 collectors of Norfolk. In July, by the kindness of the Eev. John 

 Gunn, I was enabled to see some of the matters of geological interest 

 in Norfolk, and in looking through the Yevj beautiful collection of 

 mammalian remains from the Forest-bed in the possession of 

 Mr. Jarvis, of Cromer, I observed the portion of a tooth figured 

 here. Suspecting its nature, I begged the loan of it from its 

 owner, who very courteously complied with my request. I have 

 since carefully compared it with the casts and specimens of the 

 canines of Machairodus in the British Museum, and determined it 

 as a fragment of the right upper canine of a species of that genus. 

 The only doubt which one could possibly have about such a specimen 

 as this, with its characteristic serrations and outline (see Plate XVI.), 

 was that it might belong to a large Megalosaurian whose remains 

 had been brought down from more northern Jurassic beds, and cast 

 on the Norfolk shore. 



Two eminent Paleeontologists to whom I shewed the fragment, 

 suggested this account of its origin to me, whilst two others 

 agreed in the view that it belonged to Machairodus. The 

 known teeth of Megalosaurus are, however, smaller, and of a much 

 more sharply curved outline, than is indicated by this specimen, 



