Locomotory Appendages of Trilohites. 81 



Limulm, and conseqiiently they must Lave liad ambulatory feet, 

 rather than phyllopodal feet, attached to the middle segments of the 

 body. In view of the conflict of opinions as to the nature of the 

 limbs of the Trilobites, it is to be hoped that the matter will not be 

 suffered to rest here by palceontologists, even if the most unique 

 and valuable specimens have to be sacrificed in making the requisite 

 observations." 



In 1874 Mr. S. A. Miller figured and described, in the Quarterly 

 Journal of Science, an ichnolite which he regarded as the track of 

 Asaphus. In 1880, he reviewed the work of 1874, and figured and 

 described another slab, the markings of which he regarded as made 

 by an animal generically related to the former. 



In 1875, Prof. Dana, in his Manual of G-eology, page 123, says : 

 "No remains of legs are found with any Trilobites." 



In 1876, Dr. Nicholson, in his Manual of Zoology, page 219, sa^^s : 

 " No traces of ambulatory or natatory limbs, of branchiae or of an- 

 tennte, have ever been discovei'ed. On the under surface of the 

 body nothing has hitherto been discovered except the hypostoma or 

 labrum. It has generally been supposed that the axial lobes pro- 

 tected a series of delicate respiratory feet ; but this view is doubted 

 by many authorities, and the question is one which we have at 

 present no means of deciding." 



In 1878, Prof Huxley, in his Anatomy of Invertebrate Animals, 

 page 220 (Am. Ed.), says : " Limbs or appendages capable of eifect- 

 ing locomotion are always attached either to the head or to the 

 thorax — the extinct Trilobites possibly form an exception to this rule." 

 Again, page 224 : " Now, among the water-breathing Arthropoda 

 no trace of limbs has yet been certainly discovered among the 

 Trilobites." 



In the Encyclopfedia Britannica, ninth edition (vide Crustacea), 

 Dr. Henry "Woodward, F.R.S., says: ''At present more evidence is 

 needed as to the nature of the locomotory appendages of this extinct 

 group — Trilobita." 



In a letter from Mr. C. D. Walcott, dated June, 1883, he states 

 that all his recent sections " simply corroborate the views given in 

 his pamphlet of 1881." 



These numerous references and quotations are given to show the 

 distrust and uncertainty in the minds of prominent naturalists as to 

 the limbs of the TrilolDites. 



In the autumn of 1882 a specimen of the Trilobite, Asaphus 

 megistos, was sent me for examination. In the delay of cori-e- 

 spondence with palceontologists, fortunately, no report was made, 

 for in the spring of 1883, twelve months after finding the first 

 specimen, the same party found the second, which proved to be the 

 matrix of the ventral surface of the first specimen. It was found 

 about one hundred metres from the point where the first was obtained. 



About two-thirds of the cephalic shield is broken off. That part 

 of the head antei-ior to a line drawn obliquely through the left eye 

 to the middle of the pleura of the second thoracic somite on the right 

 is entirely wanting. With the head restored, the specimen would be 



DECADE HI. — VOL. I. — NO. II. 6 



