EVOLUTION OF COLOR PATTERN IN LITHOCOLLETIS. 135 
separates Bands I and П extends entirely across the wing, but the early fusion 
of these bands on the costa is foreshadowed by the very faint tinge of buff just 
making its appearance in the costal portion of the pale fascia. On the extreme 
costa, Band V is indistinctly separated from VI. Band VI is separated from 
VII by a white streak extending entirely across the wing. With these exceptions, 
the yellow marks are defined as in the adult. The white oblique dorsal streak 
extends just within the termen, being separated from the white streak before the 
apex as in the adult. The distal end of the oblique 
dorsal streak is probably homologous with the middle 
portion of the white fascia which in primitive types 
separates Bands Vand VI. In the imago, it sometimes : 
exists as a separate small white spot before the apical ^ Fic. 19. Early stage in the 
dusting and more proximal than the oblique costal Iri gcc enc Еа Ри 
streak. Along the termen, the cilia are of an unbroken ALSO n 
yellow color, showing that at the extreme margin, there has been a complete 
fusion of Bands V, VI and VII. 
Before any of the dark margins begin to appear, the ground color deepens 
into the orange yellow of the adult. The external margin of the second fascia 
and the dark dusting at the extreme apex are considerably in advance of the 
other dark markings. The margin of the second fascia, formed by a dark line 
of seales along the inner border of Band IV, is most distinet in the middle 
portion of the wing and has not appeared on the extreme dorsal end nor on the 
eostal end beyond the angle. Its color at this stage is brownish. In one speci- 
men, the apical scales are not as dark as the margin of the second fascia; in another 
of the same age, as indicated by the state of development of the margins, the 
dusting of the apex is somewhat darker than the margin of this fascia; in both, 
however, the brownish tint is confined to the extreme apex. There is a slight 
darkening of the scales in the middle of the wing on the inner border of Band III 
adjacent to the first white fascia and along the under side of the oblique dorsal 
streak. 
The dark margins of the fasciæ and of the oblique dorsal streak continue 
to develop, but before they reach their adult condition, the scales on the outer 
side of the costal spot over vein 10 begin to darken. At this time, the apical 
dusting is still confined to the apex of the wing and is not extended along the 
termen toward the tornus. This extension of the dusting toward the tornus is a 
characteristic of specimens appearing in the summer and does not occur in forms 
from overwintering chrysalids. It develops after the other adult characters 
have appeared. 
Lithocolletis bethunella Cham. 
In the youngest specimen examined, the ground color is a uniformly pale 
ocherous, and upon it the extent of the adult markings (Fig. 84, Pl. IV) is quite 
definitely blocked out (Fig. 20). The white fascia (or second pair of streaks) 
