Wernham, The Systematic Anatotny of tlie genus Chnephora. 467 



C. Goudotii Wernham. 



I. Stern. The pith is relatively rauch bulkier than in the 

 other two species. The cells are roughly circular in section, and 

 are thick-walled throughout; no lacunae appear, and this may in- 

 dicate the absence of thin-walled tissue in the pith of the living 

 plant, in contrast with the species dealt with above. Not much 

 tannin is present in the pith; calcium Oxalate is fairly abundant, 

 a g-ood deal of it appearing as crystal-sand, but niostly, perhaps, 

 in the form of small Cluster crystals. 



The stem xylem is essentially similar to that of the other 

 two species, but the primary wood is more sharply disting'uished 

 from the pith, its elements being- considerably smaller. The fibres 

 are very thick-walled and the vessels smaller in section. The 

 wood-ring is perforated by a good many medullary rays, of the 

 type described for C. cmgusüfolia\ there is no other parenchyma 

 associated with the xylem, but a few small lacunae are to be seen 

 in the region of the primary xylem. 



The phloem consists of a narrow band almost obliterated 

 with tannin, and containing also an abundance of crystal-sand. 

 Immediately surrounding this is a fairly continuous fibrous sheath, 

 as in the other species; this comprises only a Single layer for the 

 greater part of its extent. 



The cortex is composed mostly of thick-walled cells. Tannin 

 is not so conspicuous a feature of the cell-contents as in the other 

 species, but calcium Oxalate, in the form of crystal-sand and small 

 Cluster crystals, is very abundant. This is contained mostly in 

 cells which appear to be larger than their neighbours, and in some 

 cases definite sacs, formed apparently from two or three cells, are 

 receptacles for an abundance of calcium Oxalate. 



The epidermis displays no distinctive features; no hypodermis 

 appears to be differentiated. 



II. Leaf. a) Petiole. This, unlike the two already described, 

 is approximately circular in section (Fig. 3, c). The vascular System 

 consists, as in the previous species, of one large median bündle 

 and two lateral small ones. The extremities of the former are 

 involute, and are separated by a relatively broad gap. 



The xylem of the median bündle is dense, consisting entirely 

 of vessels, which are traversed by narrow medullary rays infiltrated 

 with tannin. The phloem appears to be composed almost entirely 

 of sieve-tube tissue, appearing as a two to four-layered band ex- 

 tending continuously round the wood ring as far as the edges of 

 the gap; this tissue is packed with crystal-sand of calcium Oxalate. 

 At several points the phloem "invades" the xylem. 



As in C. angustifoUa, no definite fibrous sheath surrounds 

 the phloem; a few large fibres, isolated for the most part, appear 

 in the neighbourhood of the gap. 



The gap is in reality but one cell-layer broad, its relatively 

 greatwidth being due to the large size of the lumina of the ceUs 

 composing this layer. The tissue in the concavity of the bündle 



30* 



