Wernham, The Systematic Anatomy of the genus Canephora. 469 



are thin; and the lacunar System is more extensive than in either 

 C. madagascaricnsis or C anyustifolia. 



Calcium Oxalate is only fairly abundant in tlie lamina, but it 

 takes the form frequently of relatively large cluster-crystals. 



The central line of the transverse section is occupied by a 

 row of vascular bundles, as in the other two species; but these 

 are appreciably smaller, and associated wi J' .uwer fibres. 



The ventral epidermis is composed of cells distinctly larger 

 than those of the dorsal. In both, the cells are brick-shaped, with 

 narrower ones interspersed. The cuticles are relatively thin. 



The stomata are much more numerous than in the other 

 species, as one would reasonably expect in view of the relatively 

 mesophilous nature of this leaf. The guard-cells are not at all 

 sunk, and each is associated with a subsidiary cell slightly smaller 

 than the ordinary epidermal cells. The cell-walls are invariably 

 thin in every case. 



III. Peduncle. Unfortunately no material is available at 

 present for the purpose of anatomical investigation. 



Summary. 



A. Characters peculiar to each species. 



Each species appears to possess several anatomical 

 characters which distinguish it from the other two; these 

 are summarised below. How far these characters may be 

 constant is a question which can be decided only byfurther 

 investigation with more plentiful material. Taken collec- 

 tively, however, the characters should at least serve to 

 uphold the position as true species of the three forms 

 described, thus maintaining the conclusions based upon 

 external features. 



C. madagascariensis. I. Stem. The pith contains 

 groups of thin-walled cells. The primary xylem is asso- 

 ciated with patches of thin-walled parenchyma, which 

 may be xylem-parenchyma. Thexylemis particularly dense, 

 and there are no definite medullary rays. A regulär and 

 continuous sclerotic sheath immediately surrounds the 

 phloem. The cortex is relatively thin-walled throughout. 

 A definite hypodermis, very rieh in tannin, is differen- 

 tiated. Calcium Oxalate is, generally, less plentiful than 

 in the stems of the other two species. 



IL Leaf. a) Petiole. The xylem of the raain bündle 

 forms an almost continuous cylinder, the ventral gap 

 being extremely narrow. "Xylem-parenchyma" occurs at 

 one or two points on the ring, each group passing into 

 one of the medullary rays. The latter are conspicuous 

 and numerous, each being some three or four cells broad. 



