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intromittent organs and show the majority of species 



specific characters. For additional insight into the 

 genitalia of Coleoptera, I recommend the following: 

 Lindroth 1957; Sharp and Muir 1912; Skelley 1993; Snodgrass 

 1957; Tuxen 1970; Verhoeff 1895; Williams 1945; and Wood 

 1952. 



The median lobe is a simple tubular structure that is 

 curved and laterally flattened. The degree of curvature 

 varies from species to species and can be slightly curved 

 to arched. The shape of the median lobe's posterior end 

 varies, and can be truncate, rounded, or narrowed and then 

 rounded. 



The internal sac occasionally has pigmented areas, 

 which are patches of microspinules . These can be large 

 lightly pigmented patches of widely scattered 

 microspinules, or small dark paired patches. These patches 

 are known in only a few species. 



I found the shape of the sclerite for muscle 

 attachment at the f lagellum' s base to be important in 

 species recognition. The sclerites are all basically U- 

 shaped, but with many variations. Just posterior to this 

 sclerite is a pigmented section. This section of the 

 f lagellum is often variable in shape and curvature. A 

 close study showed it to be flexible, like cartilage. The 

 shape of this structure was not used to distinguish species 

 because it can vary from specimen to specimen. The 

 f lagellum is variable throughout the genus: long, 



