192 

 Sierra Madre Occidental), the Lesser Antilles, Central, and 



South America. 



Description . Length: 4.8-8.8 mm; Width: 2.3-3.7 mm. 

 Similar to the description under I. quadripunctatus, except 

 in the following color pattern specifics (Figs. 43-47) . 



Head black; often with orange spot. Pronotum with 

 basal black margin of variable width, often bearing 1 to 3 

 tooth-like spots; central tooth can connect with 2 free 

 central spots on disc. Each elytron has a subhumeral spot 

 variable in shape, touching or nearly touching base, 

 connected or not to scutellar spot; central band wide or 

 narrow, with edges smoothly rounded or jagged; apical spot 

 present, variable in size. Legs entirely black, or femora 

 banded orange. Genitalia dissected: 24 males, 20 females. 



Variation . Most populations in the U.S., and some in 

 northern Mexico, have the head and legs entirely black, and 

 the pronotal base narrowly margined in black. More northern 

 and central U.S. populations have the scutellar and 

 subhumeral spots separated ("Var. A" Lacordaire 1842; var. 

 antedivisa Mader 1938). Populations from southeastern U.S. 

 have these spots connected (quadripunctatus Olivier 1792; 

 var. alabamae Schaeffer 1931). This difference is clinal, 

 with the amount of spot separation variable within some 

 populations (detailed by Boyle 1954), and not deserving of 

 taxonomic rank. I refer to the U.S. color pattern, with a 

 solid black head, as "quadripunctatus" (Fig. 43). 



