196 

 Mycotretus humeralis Chevrolat (in Dejean 1836:429) is 



I. quadripunctatus (Olivier) according to Sturm (1843:305) 



and Gemminger & Harold (1876:3691). Dejean (1836:429) 



placed these names together within the same bracket, 



possibly indicating synonymy. Since no description or 



citation to a description of this name was found, it should 



be considered a nomen nudum. 



Lectotypes of both I. subcylindricus Lacordaire and I. 

 graphicus Lacordaire were chosen because there was no 

 indication that only one specimen was studied in either of 

 the original descriptions. Both designated specimens came 

 from collections apparently studied by Lacordaire (see 

 Locating Types) . 



Various authors have commented on the similarities 

 between I . quadripunctatus (Olivier) , I. graphicus 

 Lacordaire, and I. subcylindricus Lacordaire: Boyle 

 (1954:39-43), Crotch (1873b:144; 1876 : (51 ) 427) , Gorham 

 (1887:39-40; 1898:335), and Pallister (1955a:4-5). The only 

 differences observed were in the color patterns. These 

 differences proved to be clinal, as numerous intermediates 

 were found in the overlapping parts of their ranges. 



Based on the testimony of previous authors, a lack of 

 distinguishing morphological characters, the clinal nature 

 of the color patterns, and previous studies on this complex, 

 I. graphicus Lacordaire and I. subcylindricus Lacordaire are 

 synonymized under I. quadripunctatus (Olivier) . I refer to 

 these 3 as color patterns of I. q. quadripunctatus simply 



