Epwarps.—Preliminary Revision of the Crane-flies of New Zealand. 333 
Genus Macromastix Osten-Sacken. 
This genus is certainly related to Tipula, but is readily distinguishable 
therefrom by the absence of distinct verticils of hairs at the bases of the 
flagellar joints, the simple male hypopygium in which the anal segment 
is almost entirely membranous, and the fleshy and generally very short 
ovipositor of the female. These characters suggest that it may represent 
the ancestral type of the Tipulini, and in accordance with this it is 
species is such as to make it extremely doubtful whether the asinine 
Dolichopezini and Tipulini can be а as distinct. Perhaps апу 
attempt to classify the Tipulinae into sections would be es ату 
based on the male hypopygium ; in this resp ч Maer ix, Hudsonia, 
and Longurio are all similar, ды possibly might all e Я to the 
Dolichopezini, as already foreshadowed by Skuse. In classifying the 
Tipulinae a great deal 2 stress was laid by Osten-Sacken and Skuse on 
the presence or absence of a nasus at the tip of the rostrum, while 
занад ies deii poe the genus Icriomastax for a species with a 
forked n The study of the very extensive New Zealand material 
before me ee proved that these characters are absolutely worthless for 
generic separation. The nasus may be simple or forked, present or absent, 
in two obviously closely allied species, in the sexes of the same species, 
or even perhaps in individuals of one sex of the same species. I think 
it probable that Phymatopsis Skuse, Icriomastax йкы, and perhaps 
Pehlkea Enderlein, will eventually be ranked as synonyms of acromastiz. 
I zu not, however, seen specimens of any of these genera. 
may be noted further that some species of this genus have Sc, 
um. and that the fourth palpal joint is often rather short, so that the 
distinctions between the Tipulinae and Limnobiinae as given by Osten- 
Sacken are not very well maintained. A more satisfactory characterization 
of the two subfamilies is one based on the condition of the cubital vein. 
The genus Macromastix seems to be highly characteristic of the New 
to increase the number. The range of structure between the extreme 
forms is very considerable, and it is possible to recognize some well-marked 
groups of nearly allied species, ae I feel convinced that no satisfactory 
generic division can be made. most distinct groups are (a) the eight 
species placed first in the rane, н have a long praefurca (Rs) and a 
rather distinct facies; and e four species showing green coloration, 
two of which (and _— the di also) have females with rather lo ong 
and pointed oviposito Two characters which are very conspicuous in 
some species, но of the male antennae ог reduction in width of 
the cell Ax, are of specific value only and do not necessarily indicate 
affini 
Apart from the numerous New Zealand species, about a dozen species 
of Macromastiz are known from Australia, and three or four from South 
America. Walker's Tipula filigera, supposed to be from the United States, 
is also certainly а Macromastiz, the type male in the British Museum being 
in fairly good condition. 
