CunnincHaM.—The Ustilagineae, or “ Smuts,” of New Zealand. 427 
4, Tilletia Holci (Westendorp) Rostrup. (Text-fig. 16.) 
Rostr., Ust. necu p. 156, 1890. 
Polycystis Holci berg Bull. Acad. Belg., ser. 2, vol. 11, P 660, 1860. Tilletia 
Rauwenhoffii Е v. Wa ldh., учак Syst. Ust., p. 50, 187 
Sori in ovaries, piy M within the glumes, elliptical, 1-2 mm. 
long, compact, black, slightly fe 
Spores globose or subglobose | 32 x 24-28 mmm. ; epispore covered 
with a network of raised reticulations 3-4-5 mmm. high, surrounding 
Lee Pres depressions 4-7 mmm. wide, chestnut-brown. 
Host: Holcus lanatus L. tat nflorescences. Herb. No. 500. Ettrick 
(Otago), 300 m., UR B. Tennent! 10 ded 1921. 
Distribution: Europe; North / 
I have a collection of Holcus lanatus i with this species in the ovaries 
and анн striaeformis in the lea 
The three species possessing iu diste spores may readily be separated 
if the following differences are noted :— 
res under 25 mmm.  .. vci um s o Tni. 
Spores over 25 mmm 
Reticulations 2-2-5 mmm. high is s .. T. decipiens. 
Reticulations 3-4-5 mmm. high V3 : .. T. Hole. 
Furthermore, the reticulations of T. Holci are coarse, and separ. 
by large polygonal interspaces ; those of T. decipiens are closely campiei, 
being separated by much smaller areas ; whilst those of T. Tritici are 
I have been unable to germinate the spores of T. Holci. 
6. Sorosportum Rudolphi. 
Rud., Linnaea, vol. 4, p. 116, 1829. 
Sori in the form of dark-coloured pulverulent spore-masses in various 
parts of the host, chiefly in the inflorescences, formed of numerous spore- 
balls, consisting of few or many spores, at first somewhat loosely united, but 
at maturity completely separating; sterile cells a 
oloured some shade of brown, globose e ean smooth or 
verruculose ; germination similar to that of Ustilago. 
Distribution : World-wide. 
New Zealand species two—one endemic, the other indigenous; both 
are confined to the Cyperaceae. Members of e genus have xe 
been recorded on the following families: Gramineae, Cyperaceae, Јипсасеа 
Portulaceae, and Caryophyllaceae. McAlpine (1910) records thirteen s ean 
for Australia 
The genus is characterized by the rather temporary nature of the spore- 
balls, for these generally break up at maturity, when species cannot be 
distinguished from Ustilago. In one or two species the balls remain some- 
what firmly ee, when they are liable to n oe with "= sere 
"ee i 
when the spares are ek the latter by the individua seruis ey name 
firmly united in the ball by ridged folds of their epispor 
Spore-formation has been investigated by Fischer von Waldheim (1869) 
in the type species, S. Saponariae Rud. The mycelium in the infected region 
