PACKARD.] ANATOMY OF NEBALIA. 435 
pared with the mandibles of the Decapoda, especially of Mysis and other 
Schizopods,! in which a very long three-jointed palpus is developed. 
But the very long and large mandibular palpus and very weak protopo- 
dite may be set down as a diagnostic feature of the Phyllocarida. 
The 1st maxille (Pl. XXXVI, fig. 2, mzx!; fig. 5, mxz!; 5 a).—These are 
likewise singular and diagnostic features of this order, as represented 
by their structure in the Nebaliade. They consist of a small lobe (Fig. 
5 a, cv') with about 8 stout sets, and a larger lobe (cx) with the outer 
edge fringed with long coarse setz, one of which is a large ciliated seta; 
from this arises, after bending on itself at its base an extremely long and 
slender multiarticulate process (or endopodite?) which, in the female, 
is directed upward and backward (Fig. 5 a, en), reaching to the tergum 
of the basal abdominal segment, and ending in two very long slender 
sete, while a few other similar setz arise, one from each joint.? In the 
male of N. geoffroyi, according to Claus, the long setose process is 
directed forwards and downwards. : 
The 2d maxille (Pl. XXXVI, figs. 2, 5, ma”).—These are entirely unlike 
those of the first pair, and unlike the Decapodous or Phyllopod type. 
They consist of a basal portion composed of four thin, delicate, unequal 
lobes (Fig. 5, »***), edged with long setz, with two sete twice as long 
as the others arising from the 4th lobe; from this 4-lobed basal joint or 
coxopodite arise two appendages, the anterior (exopodite, ex), small, 
1-jointed; the posterior (endopodite, en), 2-jointed, the end of the second 
joint carrying above.5 long, spreading, stout, slender sete. This two- 
jointed appendage Claus considers as representing the stock of a palpus. 
This pair of maxillz are quite unlike those of Decapods (Afysis, etc.), 
as well as those of the Phyllopods, and appear to be another diagnostic 
feature of the order. 
The absence of any maxillipedes, or of any rudiments of them, either 
in the adult or in the embryo, is a negative character of a good deal of 
importance when we regard the affinities of the group to the Decapods, 
or the zoéa-form of the same order, where two (Macrura) and three (Bra- 
chyura) pairs of maxillipedes are present, there being three pairs in the 
adult Decapod. 
The eight pairs of Phyllopodiform thoracic feet (Plate XX XVII, fig.3).— 
The maxillz are directly succeeded by eight pairs of leaf-like thoracic 
feet, the maxillipedes not being present. The feet all repeat each other 
in form, and a description of the 3d or 4th pair will answer for the Ist 
as well as the last. The leg (Fig. 3, 3d or 4th pair) consists of a broad, 
thin, six-jointed appendage, the endopodite (en), which is fringed with 
very long delicate sete, those arising from the terminal joint being cil- 
iated; while a second series of fine stiff setze arise obliquely from the 
edge. To the second joint of the endopodite are appended a distal 
or lower very broad thin gill, not quite twice as long. as broad, and 
which reaches to the end of the endopodite, while situated more exter- 
nally is a double broad large lobe which corresponds to the exite or fla- 
bellum of the Phyllopod foot, this flabellum being as long as the entire 
endopodite, but not quite so broad as the gill. The distal portion of the 
flabellum is more pointed than the proximal, and, as will be seen by re- 
ferring to the figure, is more actively engaged in the process of respira- 
tion. The figure shows by the dotted lines of parenchymatous matter 
1Compare G. O. Sars. Monographi over Mysider, 1870; Pl. I, fig. 8. Claus states 
that the large palpus is very similar to that of many Amphipoda, but apparently 
overlooks the still closer resemblance to that of Mysis. 
2Claus draws attention to the position of this foot as compared with the 2d max- 
ille (putzfuss) of the Ostracoda. 
