304 



They will bavo been equally in force in the past, and must have had an influence on the phytogeny, and find 

 expression in at the present day less nearly allied species. Indeed, there is much reason for the conclusion 

 that the juvenile forms of the vegetative organs often point to relation of species apparently now very 

 distant. 



(p. 93). E. peltata Benth., is the first example, This species of an inland north-eastern district, 

 which occurs in the neighbourhood of the Burdekin, Lynd, and Gilbert Rivers, is the only species of the 

 genus which has peltate leaves in the full-grow n state (Mueller's error again repeated, see Part XLI1. p. 33. — 

 J.H.M.). Otherwise ii shows in many characters, especially in the very important shape of the fruit, a 

 great approach to E. laiifolia. It is more t han probable that the two species are closely connected ; indeed, 

 in their geographical distribution they belong to the same region, i.e., north-east Australia. Unfortunately 

 the ontogeny of E. latifolia i- not perfectly known. 1 could nowhere find a description of the juvenile 

 leaves. Generally speaking, the peltate leaf in Eucalyptus is a character of juvenile form, which 1 saw in 

 surprising perfection in E. ccdophylla and in E. erythrocorys. 



(p. 93). Eucalyptus cordata. Another case is given by .Mueller under E. cordaia. This Tasmanian 

 species remains mostly shrubby. It belongs to the species with sessile, opposite, cordate, and orbicular 

 leaves with crenate margins, possessing therefore prominently the characters of juvenile leaves. Crenate 

 leaves, says Mueller in "' Eucalyptographia, 5 ' occur also in /•.'. urnigera, and, strange as it may appear, 

 /:'. cordaia is nearest allied to this species. In its full-grown slate E. urnigera has alternate, dark green, 

 lanceolate-falcate leaves on long petioles, the calyces are elongate and narrowed at the base into distinct 

 pedicels, the operculum is larger and the fruits are urn-shaped with sunk valves, but Mr. Stephens found 

 on " OKI Man's Head," a sub-alpine mountain near Lake Crescent, trees which formed, to all appearance, 

 a perfect transition between E. urnigera and E. cordata. 



(p. 94). Further, Augustus Oldtiehl sent many years ago from Mount Wellington, near Hobart, 

 sterile specimens of the juvenile state of E. urnigera, whose lower sucker-shoots could not be distinguished 

 iu foliage in any way from E. cordata, and which had to some extent also the same whitish bloom. On 

 Mount Wellington 1 collected a state of E. urnigera whose leaves were nearly oval and whose fruits were 

 "" truncate-ovate.'* (The mature leaves of E. urnigera are alternate, petiolate, and lanceolate. See tig. 15, 

 Plate 80.— J.H.M.). 



Then follow some notes on E. ielragona and its affinity to E. eudesmioid.es, which 

 I have translated and quoted in Part XLVI, pp. 162 and 168. The affinity is explained 

 at the bottom of p. 168. 



We then pass on to Dr. Diels's " General Remarks concerning Eucalyptus," 



(p. 95). 



Tie- cases described on page 89 (E. Risdoni and E. amygdalina) enlarge the validity of the previously 

 explained connection between juvenile forms and flowering maturity, but its limits are not yet reached. 

 We clear a way for it if we try again to lay down the characters of the primary stale of the heteroblastio 

 Eucalypts. The following forms appear to be essential, or, at least, specially common in the juvenile state. 



(p. 96). 1. Shape. Leaves opposite, connate in pairs or sessile or shortly petiolate, the blade often 

 I it e at the base, occasionally even the petiole is superbasal (peltate). 



J. Bloom. Frequently all parts have a whitish or glaucous waxy bloom. 



•''. Hairs. Frequently the axis and leaves are covered with bristly hairs. 



We find that a serie- ol species of Kucalyptus produce 1 flowers in the vegetative juvenile state as well 

 as in the matured vegetative state, and i herefore develop two generative identical parallel forms, as shown 



on pages 90 and ( .i I . In other cases, 1 1 xistence of Buch pairs of identical species has been at least shown 



.i probable (pp. 93, 94). lint then' an- still other states in Eucalyptus, There are normally flowering 

 ics with juvenile leaves, of which we have not yet found their doubles with mat ure foliage. I mention 



three of the loo-t prominent species of this kind. 



1. /:'. (jamophljlla F.V.M, Leaves all opposite and broadly connate, often cordate, equal-sided, of 

 whiti .lour. An always slirubliv species inhabiting the dry north-west quarter of Australia, and 



tiding from the Hammersley Ranges Bouth-east t" tic upper Finke River. The relationship of this 

 species is doubtful. 



