

... 



... Ic 



95 





... 7c 



149 



... 



... 5c 



150 



. .. 



... Ic 



150 



.. ■ 



... 8c 



148 





H 



149 



• . . 



... U 



149 



... 



.... H 



179. 



553 



(see his figure in Eucalyptographia.) I have only seen this glandular appearance on 

 E. leucoxylon and E. Caleyi, and it should be looked for on other species. Diels figures 

 it on his E. Forrestiana." This may be seen in fig. Ic, Plate 95. I omitted to figure it 



in E. Caleyi. I have also observed it in E. megacarpa, see fig. 6b, Plate 78, and also 



~ Fig. Plate. 



E. Forrestiana 



E. eremophila 



E. macrandra 



E. occidentalis var. grandijlora ... 

 E. occidentalis 



E. occidentalis var. astringens ... 

 E. Stowardi 



and also (not figured) in filaments of E. pyropliora Benth., from near Meda, North-west 

 Australia (W. V. Fitzgerald.) It will be observed that these glandular filaments are 

 common in the Cornutse. 



Nectaries. 



To what extent the filaments of Eucalyptus assist in elaboration of nectar or 

 honey remains a subject for research. It is still more problematical what are the 

 functions of the glands on the filaments to which we have just referred. We know 

 that a filament is one of many organs that may perform the function of a nectary, and 

 so the present may be taken as a convenient opportunity for some generalisations in 

 regard to Nectaries. 



In Henslow's " Origin of Floral Structures," 141 (1888), speaking of the function 

 of nectaries in flowers, he says: ""In Violets, Atragene, Pentstemon, and Stellar ia, the 

 filaments undertake the duty," but the examples cited are not identical with what 

 obtains in the filaments of Eucalyptus. Nor does Fagopyrum, the genus quoted by 

 Goebel (" Outlines of Classification and Special Morphology," p. 381) as having 

 outgrowths or warts between the filaments come even as close to the character noted in 

 Eucalyptus. Coulter, Barnes, and Cowles, II, 844, remark that nectaries may be 

 connected with the stamens or with any other floral part. 



' They are usually very inconspicuous, and — which is very significant with 

 respect to the relation of morphology with physiology — notwithstanding their 

 enormous physiological importance, they are attached to no definite part of the plant 

 in a morphological sense, almost every part is able to perform the function of a 

 nectary." (Sach's " Text-book of Botany," p. 430.) 



The subject is discussed in the usually lucid manner of the authors in Kernel 

 and Oliver, II, 170 to 182. At p. 172 we have notes on the secretion of honey through 

 stomata; bracts (p. 173) ; fleshy disc (p. 173) ; base of ovary (p. 174) ; carpels (p. 175). 

 " Nectaries are found much more frequently on the stamens. They occur there in all 

 shapes and sizes." And so on (p. 175.) Floral leaves (p. 176); region of the corolla 

 (p. 177); nectaries interpolated between the floral leaves and the stamens (p. 178). 

 I 



