555 



of flowers of Eucalyptus have white or cream-coloured filaments; those with very 

 show}* crimson or scarlet or yellow rilanients are mostly entirely confined to western 

 and tropical Australia. In eastern Australia E. sideroxylon A. Cunn., an Ironbark, 

 very commonlv has individual trees with pink or crimson filaments. In the western 

 New South Wales Drooping Box (bicolor) the filaments are often pink, as well as white, 

 but never in anything like the abundance of sideroxylon, while in a number of species 

 such variation in coloration has also been observed, but only rarely. In Western 

 Australia there are E. erythrocorys F.v.M. and E. Preissiana Schauer, with bright yellow 

 filaments, and E. ficifolia F.v.M., E. macrocarpa Hook., E. pyriformis Turcz, E. tor quota 

 Luehmann, E. erythonema Turcz., E. pliceuicea F.v.M., and a few others have pink 

 to crimson filaments, while those of E. miniata A. Cunn. are orange-coloured. 



The nature of the pigments in flowers is known to chemists who specialise on 

 the subject, but much remains to be done. A pointer is an article, " The Colours of 

 Flowers," in " The Gardeners' Chronicle " for 29th September, 1917, p. 130. See 

 also the brief notes on Anthocyamn offered at Part LVI, p. 331. The colour of a 

 Eucalyptus flower depends almost entirely on the colour of the filaments, the 

 exceptions being the few cases of contrast of filament with anther. 



Arrangement of species according to colours is not as simple as appears at first 

 sight, because so many of them are bi-coloured, and some could, without any stretch 

 of the imagination, be considered as having at least one additional colour. The 

 following list shows all the bi-coloured species that I have seen or heard of. It is 

 probable that very many more will be found. The normal colour is white or cream, 

 except where otherwise indicated. 



1. Pink to Crimson ; Scarlet. 



E. anguhsa (E. incrassata var. angulosa). — Purple bases to filaments. See 

 Part IV. 108. 



E. bicolor are normally cream-coloured, but occasionally pink or purple. In 

 p. 312. Allan Cimningham's MS. Journal, under date 30th June, 1817, we have : " We 

 made the angle of a large deep lagoon, of considerable depth, thinly dotted with trees 

 that had marks of inundation about -4 feet above the present level of water and a few 

 inches above the general flatness of the plains. I here gathered specimens of a 

 species of Eucalyptus having a submucronated hemispherical operculum, and flowers 

 in terminal panicles of two colours (red and white), a tree of about 30 feet." 



E. Bosistoana F.v.M. — Filaments more or less suffused with purple. Marulan, 

 Xew South Wales, January. 1904. 



E. ccesia. — Pink or vieux-rose. I have also seen the colour crimson. Pink, I 

 think, is the normal colour. 



E. calophylla, — In Part XLIII, pp. 79, SO, I have discussed at some length the 

 colours of the filaments in this species and the allied E. ficifolia. They both display 

 considerable range in colour, though E. calophylla is normally white or cream, and 

 E. ficifolia is orange to scarlet. 



