12 BULLETIN 1754, U. S. DEPARTMENT OF AGRICULTURE. 
The reciprocal of ear No. 1592 is No. 1621 (Table IV). There 
was also a self-pollinated ear from the male parent of No. 1592, No. 
1622 (Table IV). The relations of these two ears are shown in 
figure 2. 
The difference between the two reciprocal ears is 5.92.62 per 
cent. The self-pollinated ear obtained from the horny parent 
does not show a significant deviation from 25 per cent. The plant 
which bore ears Nos. 1621 and 1622 is shown by the cross with waxy 
to have 60.8 per cent of the male gametes bearing the waxy charac- 
ter and 54.9 per cent of the female gametes with this character. 
The expected result of self-pollinating a plant with such a gametie 
series would be an ear with 33.4 per cent waxy, from which ear No. 
1622 with 28.5 per cent deviates 4.9+2.1, a deviation which is not 
significant. - The plant, then, which bore ears Nos. 1621 and 1622 
produced an excess of gametes bearing the waxy character, this ex- 
cess being greater in the male gametes. An excess of male gametes 
bearing the waxy character is contrary to the results obtained with 
most of the ears where it was found that the waxy gametes were 
below instead of above the expected percentage. 
The observed excess of waxy seeds on ear No. 1592 might be ex- 
plained by the assumption that a failure of the dominance of the 
horny endosperm resulted in some of the heterozygous horny seeds 
being classified as waxy. Another generation grown from the waxy 
seeds would throw hght on this matter, since if the observed devia- 
tion was due to a failure of dominance 10 of every 100 plants grown 
from the waxy seeds, when self-pollinated, would be expected to 
_ result in ears with some horny seeds, 
HORNY X WAXY. 
As the result of pollinating plants grown from the horny seeds 
with plants grown from the waxy seeds of ears Nos. 1099 and 1111, 
39 ears were obtained (Table IV). In this same table are also 
shown the ears resulting from self-pollinating the heterozygous 
horny plants. These ears will be discussed later, but it was thought 
advisable to include them in Table IV, since it makes possible the 
comparison of the behavior of the horny plants when self-pollinated 
with their behavior when pollinated with homozygous waxy. Al- 
though only 2 of the 29 ears deviated from the expected percentage 
in excess of three times the probable error, the deviation for the total 
number of seeds is 3.9 times the probable error. 
