INHERITANCE OF WAXY ENDOSPERM IN MAIZE. 41 
The percentages for the three ears are 39.3, 34.3, and 36.8 white. 
These figures are closer approximations to the Mendelian dihybrid 
proportion of 43.75 per cent than to the expected 50 per cent white. 
The nature of the crosses, however, prohibits a dihybrid ratio. 
Three ears, Nos. 1610, 1615, and 1653, were all borne on plants 
grown from white seeds and were pollinated with plants grown 
from colored seeds of the hybrid Dh 234. The plants grown from 
the white seeds could be making but one class of gametes, (7, and the 
plants grown from the colored seeds were making two classes of 
gametes, CR and Cr. 
A self-pollinated ear, No. 1703, was secured from the male parent 
of ear No. 1653. If the 39.3 per cent of white seeds on ear No. 1653 
was due to the male parent being equally deficient in both male and 
female gametes bearing one of the two color factors the expected per- 
Fed. 16/0 LE 
Fic. 4.—Diagram showing the relations of ears Nos. 1610 and 1579. 
centage of white seeds on a self-pollinated ear would be 15.45 per 
cent, but since the deficiency was observed only for the male gametes 
a percentage of 19.65 per cent could be expected. The observed pro- 
portion on ear No. 1703 was 22.3 per cent, which is not a significant 
departure from the monohybrid 25 per cent or the 19.65 per cent indi- 
cated by the deficiency of colored seeds in ear No. 1653. 
Further analysis is also possible for ear No. 1610, which had a 
reciprocal ear in No. 1579. This reciprocal ear gives a close approxi- 
mation to the expected 50 per cent, while ear No. 1610 was below 
the expected by 8.4 times the probable error. The difference be- 
tween these two reciprocals is 18.7--2.4 per cent, a difference of 7.8 
times the probable error. No analysis is possible for ear No. 1615, 
but for the two ears numbered 1610 and 1653 there is indicated either 
