INHERITANCE OF WAXY ENDOSPERM IN MAIZE. 45 
Only one ear, No. 1647, deviated by an excess of four times the 
probable error, being 8.4 per cent below the expected. 
Three ears were borne on the plant that produced ear No. 1647. 
One ear, No. 1646, was the result of self-pollination and had 21.5 
per cent white, the expected being 25 per cent. The other ear was 
pollinated by a homozygous white plant and had 46.3 per cent white. 
Although these last two ears did not deviate by a number too large 
to be bed to chance, the fact that they were both below the ex- 
pected taken in conjunction with the fact that ear No. 1647 was 
significantly lower than the expected would seem fairly conclusive 
evidence that there was a deficiency of female gametes bearing white 
aleurone on the plant that produced these three ears. 
The four groups that make up the total in Table XIX are the 
reciprocals of the four groups that make up the total in Table X VIII. 
With the exception of the first group, the families resulting from 
pollinating plants grown from heterozygous colored seeds with plants 
grown from homozygous white seeds have a higher percentage of 
white seeds than the reciprocal cross. 
The percentages for the families and the difference between the 
reciprocals are shown in Table XX. : 
TasBLE XX.—I/nheritance of aleurone color in the progeny of the two maize ears 
Nos. 1099 and 1111, by groups as shown in Table XVIII and their reciprocals 
as shown in Table XIX. 
{The minus sign (—) denotes a difference between reciprocal groups, the opposite of the remaining differ- 
ences.] 
Percentage of white seeds. 
Progeny of— Nature of cross. aes ne : 
Table X eciproca « : 
group. aaa Difference. | D +E. 
aM OS L009. 245252 b deed see WEES CHs a | 49.4+0.69 49. 0+0. 54 —0.4-+0. 87 0. 46 
LD ING 3 1 [i ee betes Gosess.2- ase 47.0+ .56 50.38+ .55 3.3+ .78 4.25 
MARIN Os O00. 22. bee seo2 cake WELDS GSS E52} 46.64 .50 50.34 .59 3.74 .78 4.77 
Loi (Pr 0 bb a Se ee eee peers GOs... 2-220 ee 45.64 .85 49.34 .95 3.741. 28 2.90 
DY GAD Sas a eee ee a Rn 47.2+ .31 49.8+ .31 2.64 .43 6.0 
The difference between the reciprocal groups of 2.6 per cent should 
be expected to occur as the result of chance but once in more than 
19,000 times. With the one exception, the groups representing crosses 
between homozygous white plants and heterozygous colored plants 
had a lower percentage of white seeds when the heterozygous colored 
plants were used as the male gametes. This would indicate a de- 
ficiency of male gametes bearing the colorless aleurone or an excess 
of the female gametes bearing colored aleurone. 
In analyzing the inheritance of waxy endosperm, a deficiency of 
male gametes bearing the waxy character was observed for the 
