BBW 



UNITED STATES DEPARTMENT OF AGRICULTURE 



BULLETIN No. 737 



Contribution from the Bureau of Entomology 

 L. O. HOWARD, Chief 



Washington, D. C. PROFESSIONAL PAPER 



March 17, 1919 



THE TOBACCO BEETLE: AN IMPORTANT 

 PEST IN TOBACCO PRODUCTS 



By 



G. A. RUNNER 



Entomological Assistant 



[With Technical Descriptions of Coleopterous larvae 

 By Adam G. BSving ] 



CONTENTS 



Page 

 A Pest In Cured and Manufactured To- 

 bacco; Its Common Names .... 1 



The Character of Its Injury 2 



Classification and Synonymy ..... 4 

 Food Substances of the Tobacco Beetle . 5 

 Food Habits of Beetles Related to the 



Tobacco Beetle 

 Losses Due to the Tobacco Beetle . 

 Distribution and Dissemination 



Origin and History 10 



Economic History . 11 



. Page 



Description of Stages 12 



Life History and Habits ...... 14 



Seasonal History . 26 



Insects Likely to be Mistaken for the 



Tobacco Beetle 27 



Natural Control 30 



Remedial Measures 37 



Preventive Measures 53 



Summary and Recommendations ... 68 



bliogrsphy 69 



