10 IV. ANONACE.E [Saccopetalum 



adnate on the outside of the broad connective. Ovaries 1-celled, as a rule 

 hairy and numerous, with the stamens on a convex or conical, rarely flat or 

 concave torus. Fruit usually consisting of numerous 1- or many-seeded carpels, 

 dry or succulent, on stalks, often long, which lengthen as the fruit ripens, 

 rarely dehiscent or united in one fleshy fruit. Seeds large, embryo small, 

 albumen ruminate, often separable into 2 halves marked by a longitudinal 

 furrow on the testa. 



Remarkable anatomical characters : — Spherical or lenticular cells containing resin in 

 the parenchyma of leaves, often visible as pellucid dots. On a transverse section the 

 timber of all species shows numerous fine transverse bars, at right angles to the 

 medullary rays, consisting of wood parenchyma. Hairs, when present, 1-celled or few- 

 celled, stellate in some genera. 



A. Ripe carpels indehiscent, distinct, mostly stalked. 

 I. — The 3 outer petals small, similar to sepals. 



Ovules6 or more, inner petals always saccate at base 1. Saccopetalum. 

 Ovules 1-2, inner petals mostly thin, flat. 



Connective acute, stamens few or numerous . 2. Miliusa. 

 Connective broad, obtuse, stamens always nu- 

 merous 3. Phjeanthus. 



II. — Petals uniform in shape and texture, fl. globose or campanulate. 

 Petals imbricate in bud, ovules 6-12 . . .4. Sager^ea. 

 Petals valvate in bud, ovules 1-2 . . . .5. Popowia. 



III. — Petals uniform, erect or spreading, the inner wanting in 2 species of Unona. 

 Pipe carpels numerous. 

 Petals valvate in bud. 



Ovules 1-2, ripe carpels 1-seeded . . .6. Polyalthia. 



Ovules 2-8, 1-seriate, ripe carpels constricted 



between seeds 7. Unona. 



Ovules many 2-seriate, connective lanceolate . Canangium (p. 16). 



Petals imbricate in bud, ovules many, 2-seriate, 



connective obtuse 8. Uvakia. 



IV. — Petals uniform, erect or spreading, ripe carpels 1-6, 



few seeded 9. Alphonsea. 



V. — All petals, or the inner only, connivent, closing over stamens and ovaries. 



1. Inner petals dissimilar, generally smaller, conniving over stamens and 



ovaries, the blades cohering by their edges. 



a. Inner petals narrowed into a linear claw. 



Stamens 6-12, ovoid, anther cells contiguous . 10. Orophea. 



Stamens co , cuneate, connective broad, truncate or 



triangular 11. Mituephora. 



b. Inner petals not narrowed into a linear claw. 



Style oblong or clavate Oxymitra (p. 19). 



Style cylindrical, generally bifid . . . .12. Goniothalamus. 



2. Petals nearly equal, similar in shape (the inner sometimes smaller), 



all or the inner only with a concave base. 



Ovaries many, petals thick, fleshy, connivent, the 



inner smaller 13. Melodorum. 



Ovaries 1 or more, petals narrow linear, from a 



concave base Xylopia (p. 20). 



Ovaries 2 or more, ovules 2, petals flat from a con- 

 cave base, fl. on woody, hooked peduncles . . 14. Artabotrys. 



Ovary 1, ovules numerous, calyx cup-shaped, 



truncate, or deeply 3-cleft Cyathocalyx (p. 21). 



B. Pipe carpels dehiscent, distinct Anaxagorea (p. 21). 



C. Carpels united into a large fleshy fruit . . . .15. Anona. 



1. SACCOPETALUM, Bennett; Fl. Brit. Ind. i. 88. 



Deciduous trees, fl. bisexual. Inner petals saccate at base, much larger 

 than sepals and the three outer petals. Stamens indefinite, connective pro- 

 longed into a conspicuous appendage. Ovaries indefinite, ovules 6 or more. 

 Species 5, 1 Australian, the others of Eastern Asia. 



