﻿Keproductive 
  organs 
  of 
  Taenia 
  pisiformis. 
  387 
  

  

  in 
  detail, 
  referring 
  to 
  it 
  merely 
  as 
  a 
  precocious 
  sjmapsis. 
  But 
  if 
  a 
  

   synapsis 
  does 
  occur 
  here 
  and 
  if 
  later 
  "the 
  spireme 
  spreads 
  thruout 
  

   the 
  nucleus 
  . 
  . 
  . 
  and 
  takes 
  on 
  the 
  appearance 
  of 
  the 
  typical 
  resting 
  

   nucleus 
  about 
  to 
  undergo 
  maturation" 
  (1. 
  c, 
  p. 
  144), 
  what 
  becomes 
  

   of 
  the 
  paired 
  chromosomes 
  during 
  this 
  intermediate 
  period? 
  Are 
  

   we 
  to 
  suppose 
  that 
  they 
  remain 
  so 
  during 
  the 
  resting 
  stage 
  

   until 
  maturation 
  sets 
  in? 
  Further, 
  what 
  is 
  the 
  meaning 
  of 
  such 
  a 
  

   phenomenon 
  ? 
  ^) 
  

  

  During 
  the 
  periods 
  of 
  oogonial 
  division 
  and 
  of 
  growth 
  and 
  skein 
  

   formation 
  there 
  is 
  a 
  marked 
  increase 
  in 
  the 
  chromatin 
  of 
  the 
  oocytes. 
  

   From 
  a 
  few 
  granules 
  in 
  the 
  young 
  oogonia 
  this 
  increases 
  until 
  

   nearly 
  the 
  entire 
  nucleus 
  is 
  filled 
  with 
  the 
  heavily 
  staining 
  skein, 
  

   which 
  opens 
  up 
  as 
  the 
  nuclei 
  increase 
  in 
  size, 
  the 
  latter 
  becoming 
  

   correspondingly 
  less 
  dense 
  (Figs. 
  33, 
  34, 
  52 
  and 
  56j. 
  The 
  greatest 
  

   increase 
  in 
  chromatin 
  occurs 
  during 
  skein 
  development, 
  not 
  during 
  

   the 
  oogonial 
  period. 
  At 
  this 
  time 
  the 
  nuclei 
  are 
  apparently 
  multi- 
  

   plying 
  too 
  rapidly 
  to 
  permit 
  of 
  any 
  great 
  chromatin 
  increase. 
  In 
  

   the 
  case 
  of 
  the 
  oocytes 
  as 
  in 
  that 
  of 
  the 
  spermatocytes 
  the 
  source 
  

   of 
  chromatin 
  increase 
  is 
  doubtful. 
  It 
  is 
  also 
  uncertain 
  whether 
  there 
  

   is 
  any 
  causal 
  relation 
  between 
  nuclear 
  and 
  cytoplasmic 
  increase 
  

   here; 
  I 
  incline 
  however 
  to 
  the 
  view 
  of 
  Richards 
  (1911) 
  that 
  the 
  

   two 
  processes 
  are 
  merely 
  synchronous, 
  not 
  interdependent.^) 
  My 
  

   reason 
  for 
  this 
  belief 
  is 
  that 
  the 
  two 
  processes 
  occur 
  more 
  or 
  less 
  

   simultaneously 
  and 
  not 
  successively 
  as 
  would 
  probably 
  be 
  the 
  case 
  

   were 
  the 
  increase 
  in 
  cytoplasm 
  determined 
  by 
  that 
  of 
  the 
  chromatin 
  

   or 
  vice 
  versa. 
  It 
  is 
  doubtless 
  true 
  that 
  the 
  two 
  conditions 
  are 
  in 
  

   some 
  way 
  correlated 
  to 
  each 
  other, 
  but 
  this 
  is 
  probably 
  indirect^ 
  

   thru 
  the 
  general 
  growth 
  stimulus 
  in 
  the 
  ovary. 
  

  

  Following 
  skein 
  development 
  and 
  cytoplasmic 
  increase 
  yolk 
  is 
  

   deposited 
  in 
  the 
  oocytes. 
  This 
  is 
  first 
  formed 
  as 
  one 
  or 
  several 
  

   small 
  spheroids, 
  which 
  increase 
  in 
  size 
  and 
  vary 
  from 
  a 
  single 
  large 
  

   mass 
  to 
  several 
  smaller 
  ones^) 
  in 
  the 
  cytoplasm, 
  and 
  is 
  apparently 
  

   formed 
  at 
  the 
  expense 
  of 
  the 
  latter 
  as 
  it 
  replaces 
  it 
  to 
  a 
  con- 
  

   siderable 
  extent 
  (Fig. 
  32). 
  Whether 
  yolk 
  formation 
  is 
  in 
  any 
  way 
  

  

  1) 
  For 
  a 
  further 
  discussion 
  of 
  these 
  questions 
  see 
  page 
  404. 
  

  

  2) 
  This 
  phenomenon 
  has 
  been 
  observed 
  in 
  Moniezia 
  by 
  Child 
  (1907, 
  1) 
  

   as 
  well 
  as 
  by 
  Richards 
  (1. 
  c). 
  

  

  3) 
  Richards 
  (1911) 
  states 
  that 
  in 
  Taenia 
  the 
  yolk 
  forms 
  but 
  a 
  

   single 
  mass 
  (except 
  in 
  early 
  stages), 
  but 
  in 
  a 
  previous 
  paper 
  (1909) 
  he 
  

   says 
  that 
  there 
  may 
  occasionally 
  be 
  more 
  than 
  one 
  of 
  these. 
  

  

  