HISTOLOGY OP THlC OVIDUCT OF THE DOMe;STIC H^N. 405 



found well out on the lips of the funnel. Figures 460 and 461 

 are both from sections of the funnel lips. The fact that there 

 is no mass of glandular tissue beneath the epithelium such as 

 occurs in other parts of the oviduct has probably led to the 

 conclusion that there were no glandular structures present. 

 However, as will be shown farther on in this paper, the glands 

 found in the funnel region are very probably homologous with 

 the larger glands occurring in other portions of the oviduct. 



Bela ('10) evidently did not find any gland cells in the funnel 

 region in the pigeon. His figures i and 2 fail to show any evi- 

 dence of gland cells, although the character of the folds shown 

 in his figure i is very similar to the condition found in the hen, 

 (cf. my figure 457)- 



As we pass posteriorly along the funnel from the mouth the 

 character of the epithelium changes in many respects. We may 

 next examine a section from near the middle of the funnel, 

 i. e., corresponding to figure 458. Figure 463 shows a portion 

 of this epithelium under medium magnification. As stated 

 before, the longitudinal ridges of the oviduct are much better 

 developed here than nearer the mouth. The secondary folds 

 of the epithelium are present as before. Collections of gland 

 cells very similar to those found nearer the mouth of the fun- 

 nel occur at the bottom of the grooves. The chief difiference 

 here is in the character of the epithelium which covers the top 

 and sides of the secondary folds. Figure 463 shows a section 

 through the middle of one of these foMs with a glandular 

 groove on either side. Portions of the adjoining folds show 

 at the edges of the figure. The similarity of the glands to 

 those described for the funnel mouth Cfig. 461) is at once 

 evident. As before short cells are arranged along the lateral 

 edges of the groove, while longer cells form a hump in the 

 center of the gland pouch. One difiference is to be noted here, 

 however. In the region of the longer gland cells, there is often 

 a double row of nuclei. The row of nuclei nearer the outer or 

 proximal end of the cells correspond in position and appear- 

 ance to the gland cells on either side of the groove. The sec- 

 ond row of nuclei lie much nearer the periphery of the cells. 

 Further examination shows that the cells in which these latter 

 nuclei occur, i. e., those nearer the periphery, are ciliated. 

 Between these ciliated cells occur the non-ciliated gland cells 

 in which nuclei lie well towards the inner end of the cell. 



