408 MAINE AGRICUIvTURAI, UXPE^RIMENT STATION. I912. 



reach their greatest development in the albumen portion where 

 they are very much more numerous and very much longer than 

 in the region under discussion. fCf. fig. 462.) 



From the above description, it is clear that the tubular glands 

 are homologous, anatomically at least, with the glandular 

 grooves in the funnel region. These collections of gland cells 

 are present even in the region of the funnel mouth.' (Cf. fig. 

 461 ) As we pass back through the funnel they are still present 

 but become somewhat better developed, larger and more 

 numerous. The tendency to pouch out at the corners becomes 

 more marked the farther one gets from the mouth. Finally at 

 the beginning of the albumen secreting region the pouching is 

 greatly increased and the first of the so-called tubular glands 

 are found. These glands will ibe discussed more in detail in 

 connection with the albumen secreting portion. Also the dis- 

 cussion of their possible function is deferred until all the ana- 

 tomical facts are in hand. 



The character of the epithelium other than that of the tubular 

 glands does not differ greatly from the epithelium in the middle 

 of the funnel. The longitudinal ridges here are broader than 

 farther forward in the funnel. This "can be seen from figure 

 453 also by comparing figure 459 with figure 458, although 

 these latter figures are drawn with different magnifications. 



The chief cause of this increase in width is found in the 

 greater space taken up by the tubular glands. For this reason 

 the secondary folds of the epithelium are forced farther apart 

 and hence do not appear so numerous. Further, the develop- 

 ment of the tubular glands give relief to growth forces in a 

 downward direction, rather than a lateral one, hence the sec- 

 ondary folds are not so clearly cut as in the earlier sections of 

 the funnel (Cf. figs. 459 and 458). 



Now, as seen from figure 453 almost as much of the epithe- 

 lial surface in the middle of the funnel is taken up by the 

 glandular grooves as by the ciliated folds. With the broadening 

 of the ridges at the beginning of the albumicn portion, the 

 glandular pouches are pushed farther apart, and hence more 

 surface is devoted to the ciliated epithelium. Considering the 

 increased glandular surface formed by the tubules it seems 

 probable that this latter increases much more rapidly than the 

 space devoted to the ciHated epithelium. 



