— 133 — 



quantity paraphyses, which take rise from small cells, bordering to 

 the cavity (fig. 51). The central cells, which all the time continue 

 to augment in size, after some time sharply differ from the sur- 

 rounding cells. They evidently here play the part of the ascogo- 

 nium, for they produce ascogonial outgrowths (fig. 52). The latter, 

 as is distinctly seen on the transversal sections, grow very strongly 

 and send out branches. As to the central cells, which have given 

 rise to the ascogenous hyphae and thus take the place of the asco- 

 gonium, we can observe their remnants even in the latest stages 

 of development of the fruit-body, wiien the perithecium has stre- 

 tched out and taken a bottle shape, and when its cavity is already 

 filled with ascogenous hyphae, young asci and paraphyses. This 

 phenomenon, which one does not notice in Podospora and Sordaria, 

 can be probably explained on account, of the thick sheath of the 

 cells replacing the ascogonium; they remain consideraljly longer 

 than in the other fungi described above. At the same time it is 

 a serious impediment for watching the cytological processes produ- 

 ' ced in the Sporormia. The fixing fluids, evidently, slowly penetrate 

 the sheath and only a small quantity of the preparations obtained 

 could be considered satisfactory in this sense. The fig. 53, shows 

 that the thick walled short cells of the future knot are at the be- 

 ginning uninucleated. The hyphae, which form these cells, have 

 generally 2 — 3 nuclei. At the stages of development of respective 

 knots, as shown on figs. 47, 48, one could not once succeed in 

 finding nuclei; only somewhat later, when in the centre of the knot 

 begins the above indicated differentiation of the larger cells, one 

 could find there 1 — 2 nuclei (fig. 49, 50). They are very small and 

 it is difficult to distinguish them in the surrounding protoplasm. Only 

 in the ascogenous hyphae, the sheath of which is much thinner, 

 the nuclei can be found much more easily. 



As so all the facts in connection with the question of fusion and 

 dividing of the nuclei of the ascogonium, have to be left out. One 

 can here only . state the fact that the development of Sporormia 

 intermedia takes quite a different course, than it does in the 

 species of Podospora and Sordaria. Here the ascogonia, which in 

 the other species are formed at the development of fruit-bodies, 

 are absent and the cells replacing the ascogonium can be recogni- 

 sed as cells forming the so called ^AVoronin hyphae", for their 



