POLLINATION AND FERTILIZATION OF FLOWERS. 227 



In another instance, figure 4, each of three flowers was 

 crossed with pollen from a different source. "The. variety used 

 was the Lorillard. Number i received pollen of the same 

 variety, number 2 was given pollen of the Currant, and number 

 3 from the Peach. As in the previous instance, there is no 

 apparent effect on the form of the fruit; and the seeds gave no 

 indication of different parentage — all were apparently typical 

 L.orillard seeds. In the offspring, the differences are marked. 

 The lines are sharply drawn between the crosses with Peach and 

 Currant, the influence of the respective male parents being very 

 evident, while the Lorillard cross is apparently unaffected by 

 either of the others ; indicating that there was no error in the 

 operation, also that there has been no transfer of influence along 

 the short peduncles."* 



In an extended series of experiments with egg plants, con- 

 ducted for three consecutive years at the Cornell University 

 and the Maine Experiment Station, the most widely varying 

 types have been crossed. In no instance, however, has there 

 appeared an immediate effect of the male parent. The little 

 Round White, when crossed with pollen from Black Pekin, dif- 

 fered in no respect from other fruits on the same plant. But 

 the offspring of this cross showed very marked variations. The 

 same facts were observed regarding several other crosses. f 



Numerous instances have been reported in which the color of 

 flowers was apparently changed by the action of foreign pollen 

 the current season. Such an instance, however, has never come 

 under the observation of the writer, though numerous crosses 

 have been made with different varieties of Tropseolum, Fuchsia, 

 Silene, Phlox, Petunia, Pelargonium, and other ornamental 

 plants. 



The statement made in the previous report upon this subject 

 still holds : "It would be unwise at the present time, to assert 

 that the directing influence of pollen does or does not as a rule 

 extend beyond the fertilization of the seed. It seems not 

 improbable that pollen from a vigorous plant may make an 



* Hop. Maine Expt. Station, 1S92. p. 39. 



t Bailey and Munson, Experiences with egg-plimts. B«L -fi. CorneU Univ. E.vp. 

 Sta., p. 14; also Rep. Maine Exp. Sta., 1S92, p. SI. 



IS • 



