138 Maine Agricultural Experiment Station. 1918. 



gives 15 solid color to 1 star. The case is not strictly mendelian 

 for in the mating of star by star one solid colored animal result- 

 ed. This animal was out of a Guernsey cow with a very small 

 star, in fact only a few hairs, and by a bull with a large star. The 

 matings of Table V point strongly to the conclusion that white 

 in the region of the udder is dominant. This dominance is not 

 strict for one solid colored bull mated to a solid colored cow 

 produced an animal with an inguinal spot. 



Table IV is typical of the behavior in inheritance of the 

 other white areas found on the neck, shoulders, rump, flanks 

 and legs. Individually considered, they all are suppressed when 

 the cross includes one pure solid colored animal (that is, the 

 above areas are recessive to solid color) for the given region. 

 This recessive quality of the hereditary units for this white is 

 not strict in any of the regions as one or two exceptions occur 

 in each case. It is conceivable that there would be an associa- 

 tion between the inheritance of the different individual spots. 

 No such correlation has, as yet, been made out. In fact, the 

 data are too limited to make any such correlations which might 

 be established, significant. 



The difficulties experienced in the explanation of the inheri- 

 tance of the Shorthorn coat color, red, white and red and white 

 are familiar to all breeders of cattle. In the study of the Roan 

 coat of this breed about the only thing which the results of Wil- 

 son, Laughlin, Wentworth, Pearson and Walthers have in com- 

 mon are exceptions which each found to the interpretations of- 

 fered by the other writers. A beginning at a solution of these 

 exceptions has been made by the excellent review of the writ- 

 ings of Storer, Wilsdorf and others on white body color by Lloyd- 

 Jones and Evvard. In this review they show that two types of 

 identical white body with colored ears exist. In the Chillingham 

 cattle this white is dominant. In the Highland cattle it is re- 

 cessive. 



The demonstration of such a difference in inheritance of 

 white as that in the above mentioned breed does not quite hit 

 the case of the Roan Shorthorn for, while the presence of these 

 two genetically different whites would complicate the results, 

 it is likely that their presence would be noted because the pat- 

 tern of each is so striking. It does remain to be shown rather 

 that the piebald cattle, like the Shorthorn, have a difference in 



