Inheritance Studies of Dairy and Beef Breeds. 139 



behavior of the separate spots which compose this piebald. A be- 

 ginning of this kind of analysis has been made by Kiesel accord- 

 ing to a review by Lang. In these experiments a solid colored 

 Limburger race was crossed to a piebald race. The first genera- 

 tion hybrids were intermediate piebald. The back cross gave 22 

 solid colored and 29 piebald. The cross to the piebald first gen- 

 eration hybrids gave out of 90 offspring, 84 piebald. Unfortu- 

 nately, no record of the exact spotting has been given, conse- 

 quently, we are left in the dark concerning any difference in 

 behavior throughout the coat. It would seem, however, that his 

 results, would fall in line with the results obtained here, where 

 each individual area is treated separately. 



Analyzed by this method, there has been shown to be a 

 marked difference in the inheritance of the individual white 

 spots throughout the animal's coat. This is, perhaps, as far as 

 we should go and is the only conclusion it is intended to empha- 

 size, but realizing that there are exceptions not yet' accounted for 

 we may say white spotting in the inguinal region is, broadly 

 speaking, dominant. The spots of the rest of the piebald pat- 

 tern are, individually considered, recessive. These enumerated 

 individually, according to the region in which they occur, are 

 white on the face (star, star snip or blaze) ; on the throat; as a 

 band across the shoulders ; as a white area on the rump at the base 

 of the tail set ; on the flanks as irregular spots ; white on the tail 

 above the switch and the white stockings on the fore feet. 



The bearing of this difference in inherited behavior on the 

 general problem is at once evident. If a red coated Shorthorn 

 should carry one of these recessive white spots we should expect 

 a small proportion of cattle produced from the random mating 

 of such an animal in the Shorthorn population would be white 

 spotted. This is what has actually been obtained in point of 

 fact. The reverse is also true that if these dominant whites- 

 are mated together, we should expect that a heterozygous mating 

 would now and then take place giving a red. The evidence 

 brought forward offers a straightforward, clear explanation 

 of the anomolous behavior of the Shorthorn coat. 



