143 
the basal cell only by one sterile cell (fig. A—D). Sporangia sitting immediately 
on the basal cell I have not observed. 
The antheridia arise at the end of peculiar narrow, cylindrical, colourless or 
feebly coloured cells given off from the apical end of the end-cells, not rarely from 
cells sitting directly on the basal cell; in fig. P such a cell is even situated directly 
on the basal cell. These androphore-cells, as they may be named, are not given 
off in the same plane as the other branches but rise more or less vertically from 
the horizontally directed cells. Usually one androphore only is given off from the 
same cell, but two androphores situated near each other also occur (fig. K). As 
mentioned below, the androphore-cells may also be situated on the carpogonia. 
The end of these androphore-cells gives rise to one or two antheridia. In the first 
case a small cell, a little longer than broad, is cut off by a transverse wall (fig. F, G, K), 
in the latter the antheridial cells are cut off by inclined walls from the end of the 
androphore-cell, leaving a little point between the two antheridia (fig. H, I); this 
point may sometimes be lengthened into a short hair-like organ. 
As to the carpogonia and cystocarps, I am sorry to say that I have not arrived 
at clearness, on account of the state of preservation of the material and perhaps 
also because these organs occurred in very small number and in insufficient stages 
of development. In particular it appeared difficult to find unquestionable tricho- 
gynes. I think however that the cell shown to the right in fig. N is really a car- 
pogonium with a spermatium attached to the trichogyne. In fig. P a cell-complex, 
probably a young cystocarp, is seen bearing an androphore and quite near to it a 
thin thread, which is perhaps a trichogyne, but no spermatium is attached to the 
latter. A similar case is shown in fig. M, where a cell bears two thin, threadlike 
organs, the one being certainly an androphore-cell, the other probably a tricho- 
gyne. The great resemblance between the androphore-cells and the trichogynes 
cause great difficulty, in particular when the antheridia are formed on the side of 
the androphore-cell. Thus the case represented in fig. L might perhaps raise some 
doubt. The resemblance between the filiform organ figured here and the andro- 
phores represented in figs. H and J might perhaps suggest that it is an androphore 
with two antheridia and prolonged point; the continuity of the protoplasmic con- 
tents in the filiform organ and that of the cell from which it is given off goes 
however to prove, that these two organs belong together, being a carpogonium, 
and that the two spermatia must have come from elsewhere and become attached 
to the trichogyne. Small round cells looking like spermatia have sometimes been 
found attached to various points on the surface of the plants (fig. A, O, Q). In the 
latter case (fig. Q) the small cell was adhering to a hyaline curved cell, the signi- 
ficance of which I do not know. 
Of stages which could be supposed to be fertilized carpogonia or cystocarps I 
have only found very few. The three-celled complex situated at the side of the 
basal cell turned upwards in fig. P I regard as a young cystocarp. A similar 
three-celled stage is shown in fig. Q at the upper side, partly hidden by an over- 
