529 
figs. 1 3). These globular bodies have 
formerly by several authors (LYNGBYE, SuHR, 
KüTziNG, SCHMITZ, DARBISHIRE (1899)) been 
considered as a parasite (Actinococcus subcu- 
taneus (LynGB.) K. Rosenv., Act. roseus 
(Svar) Kürz.), but have now turned out 
to be the much reduced sporophytic genera- 
tion of Phyllophora Brodiwi growing as a 
parasite on the gametophyte. 
The greater part of the globular 
bodies consists of nemathecial filaments 
and they can therefore be designated as 
nemathecia. They are placed either on 
particular sexual leaflets and are therefore 
often described as stipitate, or they are placed 
in great number in the undulated upper 
margin of flat frond segments (fig. 503 B). In 
the first case the fertile shoot usually remains 
short, but it may happen that it develops 
into a foliole of considerable size (fig. 502 A). 
The nemathecia can be met with at 
Fig. 511. 
Phyllophora Brodiei. A, specimen from Middelfart, 
April; radial section of young nemathecium, showing 
the outer sterile cells and the fertile ones, the latter 
connected by primary pits and partly by secondary 
pits with cells of the contiguous filaments. B, fertile 
filaments from specimen gathered in Store Belt. No- 
vember 24th with sporangia in division 625 : 1. 
all seasons; they usually arise in the spring and may early attain a considerable 
Fig. 512. 
Germlings from tetraspores of Phyllophora Brodiæi sown in the beginning of De- that some nemathecia 
cember 1925. A—B, 3/, months old, 22/3 1926. C—F, 6*/. months old, *°/, 1926. G, 7 i 
months old, /, 1926. E, optical vertical section. 4, C—F, 350:1. B, 410:1. G, 560:1. (Sporophytes) which are 
D.K.D. Vidensk. Selsk. Skr.,7. Riekke, naturvidensk. og mathem.Afd., VII, 4. 68 
size. The maximal size 
is 2 to 3.5 mm in dia- 
meter; it is reached al- 
ready in June and July, 
while the nemathecia 
are only fully developed 
in winter. The sporangia 
begin to ripen at the 
close of November, and 
ripe sporangia are met 
with in December to 
February. As the ne- 
mathecia occurring in 
winter are of different 
sizes and as nemathecia 
of considerable size are 
to be found in early 
spring, it is probable 
