The procarps arise in great numbers at the edges of the young branches of 
the flattened frond, at the inner as well as the outer edges. Their development 
has been described by Scumitz, PHırLLıps and Kyrın. Referring the reader to Kyrın’s 
paper, I shall only mention that the carpogonial branch is 3-celled and that the 
supporting cell (Tragzelle) according to Schmitz and Kyrın develops directly into an 
auxiliary cell, while Paiccips thought that the auxiliary cell was cut off from the 
supporting cell after fertilization. The first gonimoblast cell is cut off from the outer 
side of the auxiliary cell; it gives rise to several lateral cells which produce each a 
gonimolobe. The wall of the globular cystocarp consists of 3—4 cell-layers. 
The tetrasporangia are produced in particular pinnæ (stichidia) which are 
very different from the vegetative ones. They are flattened, simple and lanceolate 
or branched with nearly opposite or 
alternate, divaricate branches. The tetra- 
sporangia are included, biseriate. OLT- 
MANNS (1904, p. 661) compares them to 
INNE the stichidia in Tenioma and says that 
Fee) > | they are situated beside the central axis. 
ERO er The latter assertion, however, is not cor- 
Core c rect, for there is no axial cell-row in the 
fertile part of the stichidia, owing to the 
; ; Fig 605 AR fact that the apical cell of this organ is 
Plocamium coccineum. Young stichidia. A, 560 : 1. B—C = 
350 : 1. divided by oblique segment walls, as 
pointed out by NAGEL1. The first segments 
of the fertile pinnæ are cut off by transverse walls, but after the formation of e.g. 
4 segments by this mode of division, the apical cell divides for the rest of its 
active period by alternating oblique segment walls. The segments cut off from the 
wedge-shaped apical cell are first divided by a periclinal wall. The outer daughter 
cell divides by anticlinal cell-walls and gives rise to the marginal portion of the 
cortex or wall of the stichidium. The inner daughter cell divides by two periclinal 
cell-walls by which are cut off two peripheral cells which by anticlinal cell-divi- 
sions give rise to the cortex on the two faces of the stichidium,. while the middle- 
most cell is a young tetrasporangium. The sporangia are therefore contiguous and 
form a continuous zigzag line along the longitudinal axis of the stichidium. The 
wall of the stichidium consists of only one layer of cells. When the stichidia are 
branched, the insertion of the branches is high, the breadth of the branches is 
greatest at the base and diminishes upwards. The branches are fructiferous from 
the very base. The uppermost cells continuing the zigzag row of sporangial mother- 
cells remain sterile. 
As is well-known, the sporangia divide by three parallel walls. The primary 
nucleus is first divided into two, a transverse wall is produced in the middle of 
the cell, and the two daughter nuclei enter into the resting stage before the next 
divisions take place (fig. 604 C). 
