14 . R. A. EMEerson 
have been isolated strains that lack even this minimum of color. These 
strains furnished the original stock of type IVg. In no environment as 
yet encountered has any red or purple plant color developed in type IVg. 
BROWN, TYPE V 
The brown type was first seen in 1912, when it occurred in F2 of the cross 
purple Ia x green VIc. So far as the writer has been able to learn, brown 
plant color had not been reported previously, and he is unaware of its 
existence outside of his own cultures or of stocks grown from them. 
Seedlings and young plants of type V are wholly green. Before the 
flowering period is reached, a brown pigment begins to appear in the 
lower sheaths. At the time of flowering, the culm, the sheaths, the 
husks (Plate VI, 3), and the staminate inflorescence (Plate IV, 1 and 2) 
are brown. The anthers are usually green. The brown color extends 
to the inner husks, to the culm beneath the leaf sheaths, and to the cob 
(Plate VII, 5). That hght is not essential to the development of brown 
is shown further by the fact that the color appears under several thick- 
nesses of black paper (Plate VIII, 2). It is not uncommon to find traces 
of purple associated with the brown in the brace roots and at the base of 
the inner husks (Plate VI, 3). Abnormally developed tassels, not infre- 
quently seen on plants grown in small pots in the greenhouse, in some— 
cases show a little purple (Plate XI). The aleurone of brown plants is 
always colorless, except for xenia grains, and the pericarp is either brown, 
brownish, or colorless, never red nor cherry. Brown pericarp color of 
type V corresponds to red of types I, II, III, and IV, and brownish to 
cherry of types I and III. 
GREEN, TYPE VI 
The writer’s stock of the green type originated from a single ear obtained 
at a national corn exposition held at Omaha in 1909. The corn was 
exhibited from southern Missouri, where it is grown locally. It is a 
large dent variety, rather late in season. 
Cultures of type VIe, derived from this stock, show no plant color other 
than green at any stage of development or under any environmental 
conditions to which they have as yet been subjected (Plates IV, 3, and 
VI, 4). 
