76 -R. A. Emerson 
very poor fit being shown — z?= 24.35, P=0.0002. Evidently, green 
IVg does not give the same results in F2 of this cross as does dilute sun 
red. 
It is to be supposed, of course, that green IVg differs in some essential 
genetic way from dilute sun red, else it would not remain true green for 
generation after generation while the typical dilute sun red constantly 
produces a conspicuous amount of sun red pigment. It was therefore to 
be expected that the dilute sun red class would be deficient in F, while 
the green class would show a corresponding excess. But if the 24 green 
plants in excess of the calculated number be added to the dilute sun red 
class, that class becomes too large by eleven individuals, the excess now 
becoming almost as great as the observed deficiency. Moreover, the dilute 
purple class, it must be remembered, remains greatly deficient. If it be 
supposed that the excess of greens came about at the expense of dilute 
purples as well as of dilute sun reds, a very good fit of observation to theory 
is obtained. On redistribution of the 24 greens in excess of expectation 
to the dilute purples and dilute sun reds in the 3:1 relation usually existing 
between these classes, the corrected distribution for the six classes is as 
shown below. There are almost two chances in five that the deviations 
may be due to random sampling, P equaling 0.38. 
Dilute Dilute 
ile sum ned Brown Green Total 
Color types Purple Sun red 
~ Corrected distribu- 
GION ee ke ate 309 100 85 2H 88 74 681 
Calenlated ss: -- 287 96 96 a2 96 74 681 
Difference eae ha +22 +4 —l}i —7 —8§ 0 0 
Mere closeness of fit cannot, of course, be regarded as proof of the 
supposition on which the corrected distribution was made. But there are 
other considerations which greatly strengthen the hypothesis. In the 
case of all the F. progenies listed in table 35, it was observed that some of 
the purple plants, altho quite as strongly colored otherwise as normal 
purples, had wholly green anthers in place of the usual dark purple ones 
(Plate I, 4). Likewise some of the sun red plants had green instead of 
pink anthers. In striking contrast to this, not a single dilute purple or 
dilute sun red plant with green anthers was seen in the whole lot, the dilute 
