PLANT CouLors In MatIze rir 
purples, so far as observed, having dark purple anthers and the dilute 
sun reds pink anthers, just as in the lots considered in the first section of 
this paper. Counts of the purple and the sun red plants with different 
anther colors were made for only three of the six F: progenies (table 36), 
and for these lots not every plant was noted at the time when it was possible 
te determine the anther color positively. When some anthers have become 
dry and weathered, it is impossible to tell whether they were pink or green 
when fresh. Less difficulty is experienced with purple anthers, which 
hold their color much longer. Unfortunately, the records of the three 
F, families were not made early enough for positive identification of anther 
color of all plants. Of 162 purple plants, 117 had purple anthers and 33 
had green anthers, while 12 were not recorded. Of 50 sun red plants, 21 
had pink anthers and 12 had green anthers, with 17 not recorded. In 
these. two lots the plants with purple and pink anthers were together 
about three times as numerous as those with green anthers, thus suggest- 
ing a simple monohybrid relation between colored and green anthers. 
Working hypothesis.— If the genetic factor which is responsible for green 
anthers of purple and sun red plants be assumed to cause, in the case of 
dilute purples and dilute sun reds, not merely the anthers but the whole 
plant — leaves, sheaths, husks, glumes, stalk, and so forth — to be green, 
a satisfactory working hypothesis is afforded. The factor concerned here 
has been found to be the well-known aleurone-color factor R, or else some 
factor very closely linked with it. Some of the evidence on which this 
statement is based is presented later in this paper (pages 80,98). It may be 
pointed out in passing that the relation between anther color and aleurone 
color here noted was studied by Webber (1906) some years before the 
several aleurone-color and plant-color factors had been determined. 
Since aleurone color is not primarily concerned in the present account, 
it might be less confusing if the case were regarded as one of complete 
linkage, and if some other symbol for anther color were used and all 
reference to the R factor omitted in this paper.- Until recently there 
was nothing known of aleurone-color behavior that made necessary the 
assumption of more than the simple factor pair, Rr. The plant-color 
behavior, on the other hand, as becomes apparent later, necessitates the 
assumption of a group of multiple allelomorphs responsible in turn for 
diverse combinations of colors of leaves, sheaths, anthers, silks, and other 
plant parts. The commonest combinations in the writer’s cultures are 
