78 R. A. Emerson 
strong pink anthers with deep red silks, lighter pink anthers with red- 
dish or pinkish silks, green anthers with green silks, and so on, but there 
exist also such combinations as strong pink anthers with green silks, 
green anthers with reddish silks, and the like. Moreover, different inten- 
sities of dilute sun red in leaf blades, glumes, and other parts are sometimes 
combined with various silk-color and anther-color combinations. There 
is evidence that at least several of these combinations behave as would be 
expected if each were a definite unit allelomorphic to any one of the others. 
Perhaps the most remarkable feature of this series of allelomorphs — 
or supposed allelomorphs — is the fact that a single unit behaves as a 
dominant with respect to the color of one plant part and as a recessive 
with respect to that of another part. Thus, a combination of dominant 
pink anthers with recessive green sikks is common in the writer’s cultures. 
The wholly green plants used in the crosses here under consideration are 
recessive for green silks, anthers, glumes, sheaths, husks, and other parts, 
and dominant for colored aleurone. Since the aleurone-color symbols 
Rr have long been employed in the usual way, R as the dominant and 
ras the recessive allelomorph, this usage is adhered to in this paper. The 
effect of these factors on plant color is indicated by superscripts. Thus, 
both R’ and 7’ are dominant allelomorphs with respect to pink anthers 
and reddish silks, while both R’ and 7’ are recessive for green anthers, 
silks, and so on. In the crosses here considered it is known that 7” and 
R’ are the pair concerned. With respect to plant color, therefore, as 
contrasted with aleurone color, 7’ is dominant and R? is recessive. While 
it is realized that this usage may tend to confuse the hasty reader, the use 
of any other symbols that have so far suggested themselves would result 
in greater confusion ultimately, particularly when the interrelations of 
plant color and aleurone color are taken up. 
To return to the F, behavior of crosses of green IVg with brown, the 
following notation should express the F2 results obtained, provided the 
proposed hypothesis is tenable: 
Phenotypes Plant color Anther color 
81— ABPlr’— Ia Purple Purple 
27 —_ ABPIR™— Ig Purple Green 
27 —_ABplr’ — Ila Sun red Pink 
9— AB pl R’— IIg Sun red Green 
27 —_ Ab Plr’—IIla Dilute purple Purple 
