94 R. A. EMERSON 
occur by chance once in three trials, P equaling 0.32. One of these three — 
sun reds, when crossed with a dilute purple, Ab Pl7’, gave 71 purples © 
and 77 dilute purples, all with purple anthers, where equal numbers were 
expected. . 
Three other F2 pink-anthered sun reds produced nothing but sun red~ 
plants in F;, 228 in all (group 2, table 40). Some plants of each progeny — 
had pink and some had green anthers. Small plantings of each lot were 
made in the garden and larger plantings in the field. Anther color was 
noted in the case of the garden plants only. The records show 44 with pink 
and 16 with green anthers, a deviation from a 3:1 ratio of only 1.0 + 2.3._ 
The F, sun reds are therefore assumed to have been A A BB pl pl R’7’. 
One of these F2 plants was backcrossed to green, both of type IVg and | 
of type VIc, resulting in a total of 108 sun red plants (group 2). Altho | 
no counts were made for anther color, it was noted that the cross with 
green IVg, A b pl R’, gave both pink- and green-anthered plants, while the 
cross with green VIc, ab pl7’, gave pink anthers alone. Only two of the | 
six possible genotypes of F2 sun reds were demonstrated. | 
Only one dilute purple F2 plant was tested further (group 3, table 40). | 
From self-pollination it yielded 46 dilute purple and 9 dilute sun val 
plants, all with colored (purple or pink) anthers. The deviation from | 
a 3:1 ratio, 4.75 + 2.17, is such as might be expected by chance about 
once in seven trials, P equaling 0.14. The same F, plant when back- | 
crossed to green of types [Vg and VIc (group 3) gave 85 dilute purples and 
82 dilute sun reds where equality was expected. Evidently this F, was 
AAbbPlolr'r’. 
No F, dilute sun red or green plants were tested further. One F; 
dilute sun red, however, was found to breed true, producing an F, of 30 
pink-anthered dilute sun reds. Likewise, eight F; and Fy, greens gave 
a total of 126 green plants in the next generation. 
In so far as tests have been made, therefore, the cross of purple-anthered 
purple with green IVg has behaved as expected on the basis of the hypo- 
thetical genotype assigned to Fi, namely, A A Bb Pl pl R%7’. 
Purple Ig x green IVg.— Green-anthered purples are assumed to be 
A B PIR’, and green IVg to be A b pl R’. ~The F, genotype is therefore, 
theoretically, A A Bb Pl pl R’ R’, and F, should consist of the three 
color types purple, sun red, and green, all with green anthers. Eight 
such F; progenies are recorded in table 41, group 1. The three types 
