PLant Cotors IN MaAIzE 103 
were all, presumably, Ab PIR’. Green plants of a later generation, 
grown from these greens, when crossed with sun red plants, type Ila, gave 
64 purple-anthered purples and no other types (table 45, group 1). 
Another green crossed with dilute sun red resulted in 4 dilute purples. 
Obviously the same results would have been obtained had the green plants 
used in these crosses been a b Pl 7’, instead of A b Pl R’ as they are sup- 
posed to have been. As a matter of fact, however, one of these green 
plants had homozygous colored aleurone, and therefore must have been 
ACR. The other two greens, while they had colorless aleurone, came from 
lots known, from their 3:1 aleurone-color ratios and from crosses with 
aleurone testers, to be heterozygous for C alone, and therefore A c R. 
Moreover, the green plants from lots consisting of purples and greens in a 
3:1 relation could not have been aa, for the parents of such lots, if hetero- 
zygous for A, must have produced purples and browns rather than purples 
and greens. The green plants could therefore have been nothing other 
than A b Pl R’. 
Similarly, progenies consisting of green-anthered purples and sun reds, 
and greens, in a 9:3:4 relation, are listed in table 39, group 3. Green 
plants of these lots and their green descendants might be either A b Pl R? 
or A b pl R’, or might be heterozygous for Pl. Six such green plants were 
crossed with dilute sun reds (table 45, group 2). None of these greens 
could have been of the types discussed in the earlier section of this paper, 
namely, ab Plr’ and the like, for they were shown by appropriate tests 
(Emerson, 1918) to be A c¢R and some of them have even been used as 
C testers for aleurone color. Two of these green plants crossed with dilute 
sun reds gave dilute sun reds only, 59 in all, and are consequently regarded 
as being A b pl R’. Two others by similar crosses gave dilute purples 
and dilute sun reds in the relation 20:30, a deviation of 5.0 +2.4 from the 
expected equality from plants of the genotype A Abb Plpl R’ R’. 
Two other greens were crossed with heterozygous dilute sun reds, 
AAbbpl pl R’7’, and gave dilute purples, dilute sun reds, and greens 
in the relation 69:54:106. The theoretical distribution among these 
three classes for a total of 229 individuals, based on the assumption that 
the green parent plants were A Abb Pl pl R’ R’, is 57:57:115, a devia- 
tion that might occur by chance about. once in five trials, P equaling 0.19. 
Progenies consisting of dilute purples, dilute sun reds, and greens in a 
9:3:4 relation are listed in table 43, group 1A. Descendants of one of 
