PLANT Coors IN MAIzE 109 
Green IIIg x green VIa.— In the sections immediately preceding this, 
it has been shown that intercrosses of greens may give dilute sun reds 
(page 104), dilute purples (page 106), or sun reds (page 105) in Fy, the 
particular color type depending on the genotypes of the greens chosen 
for crossing. It remains to be shown that purple Ia can be produced by 
intercrosses of greens. A cross of green Vla, aBplr’, with green IIIg, 
Ab PIR’, should give this result, F; beng AaBb Pl pl R’r’. Such 
a cross has been made, with results as expected. 
A stock of green plants was isolated from a cross of brown V, a B Pl7’, 
with green VIc, ab plr’, and was shown, by crosses with aleurone testers 
and with dilute sun red IVa, to be type VIa, a B pl7’. Another lot of 
greens arose from a cross of purple Ig with green IVg. The purple Ig 
parent was from a lot consisting of purple Ia, purple Ig, dilute purple 
Illa, and green IIIg, coming from a cross of purple Ig with dilute purple 
Illa heterozygous for R’r’. It was therefore AABb Pl PIR’ R’. 
The green IVg plant with which it was crossed was known to be A b pl R’. 
The F; of this cross consisted, as was expected, of purples and greens only. 
The purples were type Ig and must have been heterozygous for B 6b and 
Pl pl, and the greens must have been type IIIg and heterozygous for 
Pl pl, or AAbb Pl pl R’ R’. Two of these F; greens were crossed with 
one of the greens of type VIa mentioned above. The two crosses, 9659 
and 9660, resulted as expected in purple-anthered purples, type Ia, and 
pink-anthered sun reds, type Ila, in the relation 18:20. It has been 
demonstrated, therefore, that by crossing wholly green plants of appro- 
priate genotypes it 1s possible to produce purple-anthered purples, the 
most highly colored type known, a type that is dominant to all other 
types. 
Green IITg x purple Ia.— A green plant with homozygous purple aleurone 
and belonging to a family (table 39, group 4) consisting of green-anthered 
purples and greens only, and therefore theoretically being A 6 Pl R’, 
was crossed with a purple-anthered purple, A B Plr’. A purple-anthered 
purple F,, A A Bb Pl Pl7’ R’, 5350-9, was backcrossed with green IVg 
of the genotype A 6 plr’, with the result that in the next generation 
there appeared four color types, purple-anthered purple, green-anthered 
purple, dilute purple, and green, in the relation 28:22:21:29. The 
deviations from the expected equal distribution of the 100 individuals 
were such as might occur by chance in considerably more than half of 
