PLANT Cotors In Maize 111 
2.5 + 2.1, or such as might occur by chance about twice in five trials, 
-P equaling 0.42. It is to be assumed, therefore, that the same difference 
exists between green-anthered purples and greens as between purple- 
anthered purples and dilute purples, namely, a difference with respect 
to the factor pair Bb. This in turn is merely additional evidence that 
plants which in the presence of 7” are dilute purples, A b Pl, appear as 
greens in the presence of R’ r’, which is the hypothesis under test thruout 
this section of the paper. 
Purple Ia x green-anthered dilute sun red 
A purple-anthered purple, known from appropriate aleurone-color 
tests to be R R and hence A B PI R’, was crossed with a dilute sun red 
which differed from most dilute sun reds in showing much less anthocyanic 
| pigment, particularly in early stages of growth, than is usual in plants of 
| that type, and in having little, if any, color in its anthers. The Fy’s, 
| 2975, were purple-anthered purples. 2 was expected to show the four 
_ color types, purple, sun red, dilute purple, and dilute sun red, commonly 
found in crosses of purple Ia with dilute sun red IVa. As a matter of 
fact, the single F2 progeny grown was found to consist of these four color 
types as major classes, but each class was found to have colored-anthered 
(purple or pink) and green-anthered subclasses. The difference between 
the two subclasses for purple and sun red was sharp, just as is the case 
in crosses of purple Ia with green IVg, but it was often difficult. to separate 
green-anthered dilute purples from green-anthered dilute sun _ reds. 
_ Ordinarily, anther color (purple or pink) is the surest means of distinguish- 
ing between dilute purple and dilute sun red. When both have green 
anthers the separation must be based on the relative amount of pigment 
in other plant parts — a difference that is usually not very marked until 
late in the life of the plants, when dilute purples usually show materially 
-more pigment, especially in parts not exposed to the sun, than do dilute 
sun reds. It will be recalled that in crosses of purple Ia with green IVg, 
both colored and green-anthered purples and sun reds appear, but that 
all the dilute purples and dilute sun reds have colored anthers, the green- 
anthered individuals appearing as wholly green in all plant parts except 
perhaps the pericarp. But in the cross here considered, no wholly green 
plants were found. 
