114 R. A. EMERSON 
there was distinctly more red color in the leaves of the plants from colored 
seeds than in the leaves of the plants from white seeds. Particular atten- 
tion has not been given to a possible effect of the C factor on mature plant 
colors of other color types. Many cultures of dilute sun reds and greens 
have afforded opportunities for observing any effect of C and ¢ on red color 
in the leaves of seedlings, but no effects have been noted. No particular 
attention was paid to the matter at the time when the seedlings were under 
observation, but if the C ¢ pair had exerted any marked influence it would 
probably have been noted. 
Numerous cultures of dilute sun red seedlings have been noted with 
respect to possible effects of the aleurone-factor pair Pr pr, but no effect 
has been observed, the purple and the red seeds having produced seedlings 
with apparently the same intensity of red color. Likewise, no influence of 
Pr pr on mature plant color has ever been observed in the case of either 
sun red or diiute sun red. With purple and dilute purple plants, however, 
a distinct effect is noticeable. Purple and dilute purple plants from seeds 
with purple aleurone have purple anthers, while those from seeds with 
red aleurone have reddish purple anthers (Plate I, 1 and 3, and Plate 
II, 1 and 3). A similar effect is often seen also in the color of the inner 
husks. In neither the anthers nor the husks is the effect always suffi- 
ciently distinct to make possible an accurate separation of plants from 
purpleeand from red seeds if they are growing in mixed cultures. In 
some cases, however, the difference is very distinct. And when the 
seeds are separated with respect to purple and red aleurone, the two lots 
of plants resulting usually show fairly distinct differences in anther color 
and often in husk color as well. 
EXPRESSION OF PLANT-COLOR AND ALEURONE-COLOR FACTORS 
The mode of expression of the several plant-color factors has been dis- 
cussed in detail in this paper, and similar discussions of aleurone-color 
factors are available in numerous other papers. Since aleurone colors 
and certain plant colors —the purple-red series — are doubtless antho- 
cyanins, it seems natural to expect close interrelations between them. 
Many such relations have been noted in this account. There are certain 
matters, however, which need to be brought together in a summary 
discussion. 
