PLANT Coors In Maize 2 117 
Fl pl is perhaps expressed most definitely in the color of anthers, tho 
the expression is by no means limited to them. Purple (A B Pl) and 
dilute purple (A b Pl) differ from sun red (A B pl) and dilute sun red 
(A b pl), not merely in having purple rather than pink anthers, but also 
in the coloration of their inner husks, their culms, and the like. What 
little brown color is seen in a b Pl is limited almost wholly to the staminate 
inflorescence. The staminate inflorescence of purples, A B Pl, and of 
browns, a B Pl, is strongly colored, but that of dilute purple, A b Pl, 
except for anther color, is not very different from what is seen in dilute 
sun red, Abpl. The Pl factor, when associated with ro", is expressed 
in the pericarp as cherry in purple and in dilute purple, and as brownish 
in brown and in green of the genotype ab PI. 
Factors Bb and Pl pl are not known to be concerned with aleurone 
color. All the other factors affecting plant color are aleurone-color 
factors also. Of these the pair Pr pr influences anther color of purple 
and dilute purple, and to some degree the husk color as well. The pair 
Cc has been observed to affect the leaf color of mature plants of the 
sun red type. The pair A a is expressed to some degree in all such parts 
as culms, sheaths, husks, glumes, anthers, and silks. The pericarp, 
if a pericarp factor P is present, is red with A and brown with a, or if r” 
and PI are present, it is cherry with A and brownish with a. The R 
series of factors influences many plant parts. With duplex R’ or r’, 
no color develops in any part of the plant, except the aleurone, provided 
B is absent. With B these factors give colorless anthers and silks merely. 
Factors R’ and 7’, if A also is present, affect practically all plant parts 
in which anthocyanic pigments ever develop, but are not |known to 
have any influence on the color of the pericarp. The factor r” is, how- 
ever, concerned with pericarp color provided PI also is present. This 
factor has a marked influence on the amount of color that forms in the 
leaves, particularly of dilute purple and dilute sun red. 
It is of no little interest that the R series of factors, which’ behaves 
as a group of multiple allelomorphs with regard to plant color, usually 
acts as a simple pair in respect to aleurone color.6 Moreover, some of 
these factors act as dominants with respect to aleurone color and as 
recessives with respect to plant color, while the dominance of others is 
6 There isTsome evidence that at least one aleurone-color pattern is dependent on an allelomorph of 
Rr, the three thus constituting a group of triple allelomorphs affecting aleurone-color development. 
