360 University of California Piiblications in Zoology [Vol. 13 



29, fig. 67a). Proximal barbules differ from those of remiges in 

 base being relatively slender, with inconspicuous ventral teeth, the 

 pennulum being extremely slender and filamentous, nearly as long 

 as base. 



Breast feathers, as exemplified by Melopelia asiatica, with both 

 kinds of barbules characterized by elongation of bases, which are 

 relatively more slender than in back feathers, and very great 

 elongation of the slender threadlike pennula. Distal barbules (pi. 

 29, fig. 666) have slender base, all barbicels reduced, and pennu- 

 lum two or three times length of base on more proximal part of 

 vanule, shorter than base on distal part of vanule. Proximal bar- 

 bules (pi, 29, fig. 66c) with similarly elongated base and extremely 

 long pennulum, several times length of base on proximal part of 

 vanule, about equal to it on distal part, 



h) Other Types 



In the majority of the Columbidae the remiges have a structure 

 very closely similar to that described above for Columba livia. A 

 very unusual and probably recent departure from the ordinary 

 type is to be found in the forked primaries of Drepanoptila, in 

 which the entire feather plate is double on the terminal portion. 



Goura coronata differs widely from the other Columbidae in 

 the character of the proximal dorsal cilia of the distal barbules of 

 the inner vane (pi. 29, fig. 70a), which are large and specialized 

 as stout, lobelike projections as in herons and vultures. In other 

 respects the distal barbules, and in all respects the proximal bar- 

 bules, resemble those of typical Columbidae. The coverts, scapulars, 

 and back feathers undergo very little modification in the different 

 genera except for production of color, (See plate 29, figure QQa, 

 covert from Melopelia; figure 68ft, covert from Macropygia; figure 

 69a, covert from Osmotreron). The breast feathers also differ but 

 slightly in different groups. Even in Goura coronata, which has 

 feathers differing most widely from the type of any species in the 

 family, the breast feathers are remarkably similar to those of other 

 columbids (pi. 29, figs. 70c, 70tZ). The barbules on the loose-vaned 

 crown feathers of Goura (pi, 29, fig, 70e) have lost their pennula 

 and the bases are cigar-shaped. 



In the Pteroclidae, exemplified by Pteroclis arenarius, the distal 

 and proximal barbules of the inner vane (pi. 29, fig. 71fl, 71c) are 

 distinctly of typical columbid type, but those of the outer vane 

 differ in having very short pennular cells with long, slender ventral 

 cilia and very short, knoblike dorsal cilia. The scapular feathers 

 have the peculiarities of the pennulum of the distal barbules still 



