89 [Stowell. 
The Glosso-pharyngeal Nerve in the Domestic Cat. 
By T. B. Stowell, A.M, Ph.D. 
(Read before the American Philosophical Society, March 2, 1888.) 
Introduction.—The following study in comparative neurology is a con- 
tinuation of the contributions made by the author and already published 
in the Proceedings of the Society. [The Vagus Nerve in the Domestic 
Cat (Felis domestica), read July 15, 1881 ; The Trigeminus Nerve in the 
Domestic Cat, read May 21, 1886 ; The Facial Nerve in the Domestic Cat, 
read Noy. 5, 1886.] The object of these contributions has been presented 
in the previous papers. 
Iam not aware that any one has published the detailed distribution and 
relations of this nerve ; this study is therefore offered as-a contribution to 
comparative neurology. 
Preparation.—As previously recommended, the cats were injected with 
the ‘‘starch injection mass.’”” The dissections have been verified upon 
alcoholic and upon recent specimens ; the nerves have been traced under 
a magnifying power of 15-25 diameters. It is hoped that such a degree 
of accuracy has been attained as will render this study serviceable to 
students of the nervous system. 
DESCRIPTION. 
Synonymy.—Nervus glosso-pharyngeus, N. pharyngo-glossus, Glosso- 
pharyngeal nerve, Highth pair of cranial nerves (Willis), Ninth pair (Som- 
mering). 
General Characters.—The N. glosso-pharyngeus is the cephalic member 
of a group of nerves whose ectal origin is along a line caudad of the 
eminentia auditoria and dorsad of the area elliptica. The common niduses 
in man from which the ental fibres of these nerves arise have been desig- 
nated the “‘nuclei of the lateral mixed system ’’ (28, p. 747). 
The N. glosso-pharyngeus is characterized by the presence of two ganglia 
upon the central portion of the nerve-trunk, the central one (Fig. Ebr.) 
is within the foramen of exit (Fm. jugulare), the peripheral one (Fig. Pe.) 
is about 7 mm. peripherad of the foramen. The presence of the root gang- 
lion and the ental origin ally this nerve and the dorsal roots of the myelic 
nerves. 
The glosso-pharyngeus is specifically the sensory nerve of the pharynx, 
of the mucosa of the tympanum, and of the basal (circumvallate ?) papillz 
of the tongue. It includes motor fibres which are given to the cephalic 
pharyngeal constrictor and to the stylo-pharyngeus muscles. Communi- 
cating rami connect the N. glosso-pharyngeus with the N. vagus, the N. 
sympathicus, the G. oticum, and with the N. facialis. 
The glosso-pharyngeus is peculiarly the nerve of deglutition as well as 
the nerve of taste. The importance of having these functions associated 
PROC. AMER. PHILOS. SOC. xxv. 127. L. PRINTED APRIL 4, 1888. 
