208 MEMOIRS OF THE CARNEGIE MUSEUM 
sagittal crest prominent; brain-case small ; incisors heavy and short; canines compara- 
tively small and oval in cross-section ; P' with antero-internal cusp of moderately large 
size and located relatively close to the main body of the teeth; M' and M? large and broad; 
M* present, though small, practically one-rooted and aligned with the internal border of 
M* and M’. To the above cited characters should be added: 12 01 P4 M3; the 
symphysis of the inferior ramus separated or slightly codssified ; the tubercles of all the 
molar-premolar teeth blunt ; the heel of the lower carnassial tooth large: metatarsals not 
appressed at their proximal ends ; feet sub-digitigrade ; tail long and heavy. 
THE SKULL. 
(Plates LX XIV-LX XVII.) 
In the articles cited above attention was paid to the cranial and dental charac- 
ters of Daphenodon superbus. In the present paper, therefore, I will only give a 
short review of my previous description and add characters which I regard as of 
interest and importance to the student. 
The cranium of this new genus represents an animal very nearly as large as a 
fully adult gray wolf, but the skull is proportionally shorter and broader, the brain- 
case smaller, the occipital condyles smaller, the basioccipital and basisphenoid of 
greater transverse diameter, the paroccipital process further separated from the 
tympanic bulla and directed more backward, the mastoid larger, the tympanic bulla 
smaller, and the postglenoid process heavier. When compared with Daphenus 
felinus the skull is distinctly shorter and broader but the brain-case is evidently 
not much enlarged and there is a large frontal sinus; the muzzle is heavier; 
the pterygoids are shorter, and the region back of the pterygoids is apparently 
also somewhat shortened. The postglenoid and paroccipital processes are of the 
same relative size and position while the mastoid process is a truncated rounded 
knob, and is of proportionally greater size. In fact the entire mastoid region of 
Dapheenodon seems to have increased, which is a distinct step away from the line of 
Canis. The space between the paroccipital process and the condyles is shorter, so 
that the occipital condyles are more sessile in the specimen under description than 
in the Oligocene genus.? ‘The tympanic bulle are present on both sides in the type 
specimen ; they are of moderately large size and triangular in general outline. The 
tubular process, which encloses the external auditory meatus is quite prominent and 
is protected posteriorly by the heavy mastoid, though not so completely codssified 
with the latter bone as in the recent dogs. On the postero-internal angle of the 
bulla there is a sharp vertical ridge which unites with a similar though much 
®In Daphenus felinus (No, 492, C. M. Cat. Vert. Foss.) the base of the skull in the region of the condyles is some- 
what injured and the separation of the condyle from the paroccipital process may, in part, be due to this fact. 
