ZAZ MEMOIRS OF THE CARNEGIE MUSEUM 
has a simple crown; it is separated from the canine and P* by short diastemata. 
The latter tooth is also isolated, but the diastema back of it is very short. This 
tooth has two roots; its crown is comparatively simple, there being no anterior or 
posterior basal cusps. P# is simple-crowned, like the anterior premolar, and other- 
wise has about the same relative proportions as in Daphanus felinus. Its position 
in the alveolar border is oblique, which makes it appear of small antero-posterior 
diameter in the side view of the skull. In P4 (carnassial) a slight change from what 
is observed in the Oligocene genus has taken place. The deuterocone is slightly 
reduced in size and is closer to the main body of the tooth than in either Daphanus 
or any of the allied John Day forms, and in this respect is more like the corre- 
sponding tooth in Amphicyon major of Europe. The deuterocone is, however, sup- 
ported by a much heavier root than either in Amphicyon or Canis and its oblique 
position in the alveolar border is totally unlike the latter genera. The smaller 
antero-posterior diameter and otherwise primitive features of the tooth places the 
genus rather closer to the Oligocene form and removes it further from Canis. M+ and 
M® are larger than in any of the allied John Day forms (Temnocyon, Mesocyon, Para- 
daphenus) and have also undergone some changes from the corresponding teeth in 
the Oligocene genus, while M* is apparently the same. This change of the molars 
is principally due to the greater development of the postero-internal angle, which 
gives these teeth a slightly greater antero-posterior diameter internally than is 
usually seen in the Oligocene species, and the tooth is otherwise apparently inter- 
mediate between that of these early forms and Canis occidentalis, which has a very 
prominent, sharp, and crescentic postero-internal tubercle, especially on M+. The 
development of the postero-internal angle is unusually like what we find in Am- 
phicyon major and it is quite curious to find the conical structure of the different 
tubercles so nearly alike in the two genera. M* and M* of the European genus 
are, however, proportionally larger, the posterior intermediate cusp (metaconule), 
especially on Mz, is more strongly developed and the antero-external cusp (para- 
cone) is the larger, while in Daphenodon para- and metacones are subequal and of 
the primitive type met with in Daphonus ; in fact the tubercles are more depressed 
than in the latter genus, as is seen by referring to the illustrations, Pl. LXXV. The 
internal border of the grinding surface of M* extends slightly below that of the 
preceding tooth. This character, which is well shown on Pl. LX. XIV, may, in part 
at least, be due to slight displacement. M® is, as stated, practically a duplication of 
that tooth in Daphenus, i. ¢., a small tooth with transversely oval and very low 
crown with a very short root, the alveolus which is posteriorly formed by a thin and 
very delicate border in the maxillary. 
