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248 MEMOIRS OF THE CARNEGIE MUSEUM 
greater developed eminence on the lower half of the palmar face in Daphcnodon. 
The free end of the tuber is suddenly expanded, especially laterally, and there is a 
well defined groove for the tendon of Achilles, as in Daphenus and the recent dogs and 
cats generally. There is on the outer angle of the dorsal border a long and rugose 
groove, which, however, is not so deep as 
that in the Oligocene genus. This groove 
is also quite conspicuous in the recent 
lion, while in Canis it is entirely wanting. 
The articular facets for the astragalus are 
separated as usual by the groove for the 
interosseous ligament, the external facet 
being quite oblique and more gently con- 
vex antero-posteriorly than in Canis, 
while the sustentaculum is more promi- 
nent, the internal facet being slightly con- 
cave infero-superiorly and carried well 
Fig. 44. Anterior and Fibular Views of Caleaneum of over the superior edge, not unlike what is 
hire Ripe nt seen in Canis. The greater process of the 
caleaneum is much abbreviated, so that the head of the astragalus extends well be- 
low the distal end, as in Daphewnus, also observed by Mr. Hatcher (/. ¢., p. 91). The 
peroneal tubercle is well developed, causing the fibular face of the bone to be quite 
concaye supero-inferiorly. The distal end has a large facet for the cuboid, which is 
gently concave in all directions and pear-shaped in its general outline. 
The astragalus and caleaneum of Amphicyon major are represented in our col- 
lection by accurate casts, which display well marked differences from the present 
genus. In the first place the astragalus in the European form is broader and shorter 
in proportion ; secondly, the head is more sessile, due to the shorter neck ; thirdly, 
the suleus which separates the articular facet for the tibia on the tibial face of the 
inner condyle in Daphenodon is absent in Amphicyon; and fourthly, the external 
articular facet for the calcaneum is less concave supero-inferiorly than in the Amer- 
ican form. 
The tuber of the calcaneum in Amphicyon is proportionally shorter, and the 
palmar border is much heavier than the dorsal, while in Daphanodon the two 
borders are subequal in thickness, as is also the casein Daphanus. In the European 
form there is no well defined groove for the tendon of Achilles and the free end of the 
tuber calcis is much enlarged, as in the bears. The sustentaculum is prominent, as 
in Daphanodon, but the greater process of the caleaneum is more abbreviated. 
