2G0 
REV. J. T. GULICK ON DIVERGENT EVOLUTION 
of offspring belonging to tbe one variety to which the four varia- 
tions alike belong ? Evidently the variation represented by the 
fraction T 8 ^ will have the greatest influence on the following 
generation. But as the supposed conditions allow of exact com- 
putation, let us look at the problem a little closer. If each varia- 
tion numbers say a thousand individuals, then the number of 
each that will breed true will be as follows : — Of the one repre- 
sented by t 9 0 , 526 will breed true and 474 will cross, 
And the next generation of each kind will be as follows : multi- 
plying the pure parents by 10, and the hybrid parents by 8 or 9, 
according to the value of m, we have of those represented by 
There can, therefore, be no doubt that under such conditions the 
average Prepotential Segregation and Segregate Eecundity of 
the next generation will be considerably advanced, and so with 
each successive generation till the average of the Pure forms is 
represented by the fraction -fy, and is surrounded by a circle of 
variations, of which one will be represented by the fraction 1 2 fr . 
And from this new point continuous advance will be made toward 
ever higher and higher grades of Segregation and Segregate 
Eecundity ; though of course the process will be subject to 
antagonisms and limitations arising from the principles of Self- 
accumulating Vigour and Self-accumulating Adaptation. Let it, 
however, be carefully noted that we have in this process the 
manifestation of a new principle, for it rests not only on Self- 
accumulating Positive Segregation but on Self-accumulating 
Segregate Eecundity. 
450 
444-5 
400 
T 9 0 -, pure offspring 5260, hybrids 4266, 
5500, ,, 4050, 
5555, „ 3556, 
6000, „ 3200. 
